About the Authors

Zhifeng Li

Contributed equally to this work with: Zhifeng Li, Changjun Bao

Affiliations Nanjing University Medical School, Nanjing, China, Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Changjun Bao

Contributed equally to this work with: Zhifeng Li, Changjun Bao

Affiliation Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Jianli Hu

Affiliation Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Wendong Liu

Affiliation Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Xiaochen Wang

Affiliation Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Lei Zhang

Affiliation Zhejiang Provincial Center for Disease Prevention and Control, Hangzhou, China

Zhengmin Ji

Affiliation Jiangning Center for Disease Prevention and Control, Jiangning, China

Zhi Feng

Affiliation Jiangning Center for Disease Prevention and Control, Jiangning, China

Luxun Li

Affiliation Lishui Center for Disease Prevention and Control, Lishui, China

Aihua Shen

Affiliation Lishui Center for Disease Prevention and Control, Lishui, China

Xuejian Liu

Affiliation Xuyi Center for Disease Prevention and Control, Xuyi, China

Hongjun Zhao

Affiliation Xuyi Center for Disease Prevention and Control, Xuyi, China

Wenwen Tan

Affiliation Yixing Center for Disease Prevention and Control, Yixing, China

Jiangang Zhou

Affiliation Yixing Center for Disease Prevention and Control, Yixing, China

Xian Qi

Affiliation Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Yefei Zhu

Affiliation Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Fenyang Tang

Affiliation Jiangsu Provincial Center for Disease Prevention and Control, Nanjing, China

Carol J. Cardona

Affiliation College of Veterinary Medicine, University of Minnesota at Twin Cities, Saint Paul, Minnesota, United States of America

Zheng Xing

zxing@umn.edu

Affiliations Nanjing University Medical School, Nanjing, China, College of Veterinary Medicine, University of Minnesota at Twin Cities, Saint Paul, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZL CB ZX. Performed the experiments: ZL CB JH WL XW LZ ZJ ZF LL AS XL HZ WT JZ XQ YZ FT. Analyzed the data: ZL CB CJC ZX. Contributed reagents/materials/analysis tools: JH WL XW LZ ZJ ZF LL AS XL HZ WT JZ XQ YZ FT. Wrote the paper: ZL CB CJC ZX.