About the Authors

Libo Su

Affiliations College of Veterinary Medicine, Jilin University, Changchun, Jilin, China, Mucosal Immunology and Biology Research Center, Massachusetts General Hospital and Harvard Medical School, Charlestown, Massachusetts, United States of America

Yujuan Qi

Affiliations Mucosal Immunology and Biology Research Center, Massachusetts General Hospital and Harvard Medical School, Charlestown, Massachusetts, United States of America, Qinghai University Medical School, Xining, Qinghai, China

Mei Zhang

Affiliation Mucosal Immunology and Biology Research Center, Massachusetts General Hospital and Harvard Medical School, Charlestown, Massachusetts, United States of America

Meiqian Weng

Affiliation Mucosal Immunology and Biology Research Center, Massachusetts General Hospital and Harvard Medical School, Charlestown, Massachusetts, United States of America

Xichen Zhang

Affiliation College of Veterinary Medicine, Jilin University, Changchun, Jilin, China

Chienwen Su

Affiliation Mucosal Immunology and Biology Research Center, Massachusetts General Hospital and Harvard Medical School, Charlestown, Massachusetts, United States of America

Hai Ning Shi

shiha@helix.mgh.harvard.edu

Affiliation Mucosal Immunology and Biology Research Center, Massachusetts General Hospital and Harvard Medical School, Charlestown, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CS XZ HNS. Performed the experiments: LS YQ MZ MW CS. Analyzed the data: LS CS HNS. Wrote the paper: HNS.