About the Authors

Ria R. Ghai

Affiliation Department of Biology, McGill University, Montreal, Quebec, Canada

Colin A. Chapman

Affiliations Department of Anthropology and McGill School of Environment, Montreal, Quebec, Canada, and Wildlife Conservation Society, Bronx, New York, New York, United States of America, Makerere University Biological Field Station, Fort Portal, Uganda

Patrick A. Omeja

Affiliation Makerere University Biological Field Station, Fort Portal, Uganda

T. Jonathan Davies

Affiliation Department of Biology, McGill University, Montreal, Quebec, Canada

Tony L. Goldberg

tgoldberg@vetmed.wisc.edu

Affiliations Makerere University Biological Field Station, Fort Portal, Uganda, Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RRG CAC TJD TLG. Performed the experiments: RRG CAC PAO TLG. Analyzed the data: RRG TJD TLG. Wrote the paper: RRG TJD TLG.