About the Authors

Christopher Reed

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Kenny Lin

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Catherine Wilhelmsen

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Brian Friedrich

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Aysegul Nalca

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Ashley Keeney

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Ginger Donnelly

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Joshua Shamblin

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Lisa E. Hensley

Current address: U.S. Food and Drug Administration, Silver Spring, Maryland, United States of America.

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Gene Olinger

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Darci R. Smith

darci.smith@colostate.edu

Current address: Colorado State University, Fort Collins, Colorado, United States of America.

Affiliation United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LEH AN DRS. Performed the experiments: CR KL BF AK GD JS DRS. Analyzed the data: CR KL CW AN AK GD JS GO DRS. Wrote the paper: CR DRS.