There are errors in Equation (3). In the description of Equation (3), the equilibrium frequencies PIX are incorrectly defined in terms of pi, where in fact they should be defined in terms of phi. Please see the corrected description of Equation (3) here.
Reference
Citation: The PLOS ONE Staff (2014) Correction: Correcting the Bias of Empirical Frequency Parameter Estimators in Codon Models. PLoS ONE 9(8): e106200. https://doi.org/10.1371/journal.pone.0106200
Published: August 19, 2014
Copyright: © 2014 The PLOS ONE Staff. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.