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Table 1.

Bacterial strains and plasmids used in this study.

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Table 2.

Primers used for dnaK deletion and complementation.

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Fig 1.

Construction of the ∆dnaK mutant and assessment of bacterial growth under aerobic conditions.

(A) Schematic representation of the in-frame deletion strategy used to generate the ∆dnaK mutant of Staphylococcus xylosus. Approximately 1-kb regions flanking the dnaK coding sequence were cloned into the temperature-sensitive shuttle vector pIMAY for allelic exchange. (B) Growth kinetics of wild-type, ∆dnaK, and ∆dnaK-C strains cultured in TSB at 37 °C with shaking. OD₆₀₀ measurements and CFU enumeration were performed at indicated time points. Data represent means ± SEM from three independent experiments. Statistical comparisons among groups were analyzed by one-way ANOVA followed by Tukey’s post hoc test. ns, not significant.

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Fig 2.

Assessment of host cell adhesion, invasion, and intracellular survival.

(A) Quantification of bacterial adhesion to breast cancer cells at 2 h post-infection. (B) Quantification of bacterial invasion into breast cancer cells at 2 h post-infection. Salmonella enterica serovar Typhimurium was used as a positive control for adhesion and invasion, while E. coli DH5α served as a negative control. For both adhesion and invasion assays, Salmonella enterica serovar Typhimurium was used as a positive control, and Escherichia coli DH5α served as a negative control. (C) Quantification of total bacterial load per well at 24 h post-infection, determined by S. xylosus-specific qPCR. (D) Enumeration of intracellular CFUs of wild-type, ∆dnaK, and ∆dnaK-C strains at 24 h post-infection following gentamicin treatment and host-cell lysis. Data represent means ± SEM from three independent experiments performed in triplicate. Statistical significance was determined by one-way ANOVA followed by Tukey’s multiple-comparison test (p < 0.05; ns, not significant).

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Fig 3.

DnaK is dispensable for resistance to hypoxic stress and antimicrobial peptide exposure.

(A) Growth of wild-type, ∆dnaK, and ∆dnaK-C strains under hypoxic conditions (1% O₂). OD₆₀₀ measurements and CFU enumeration were performed at indicated time points. (B) Survival of strains following treatment with LL-37 (10 µg/mL, 2 h). Viable CFUs were determined by plating. Data represent means ± SEM from three independent experiments. Differences among groups were analyzed by one-way ANOVA. p < 0.05; ns, not significant.

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Fig 4.

Sensitivity of the dnaK-deficient strain to oxidative and acid stress.

(A) Survival of strains following exposure to 5 mM hydrogen peroxide for 1 h. (B) Survival after incubation in TSB adjusted to pH 5.5 for 2 h. Viability was quantified by CFU plating. Data are presented as means ± SEM from three biological replicates. Group comparisons were evaluated using one-way ANOVA with Tukey’s multiple-comparison test. p < 0.05.

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Fig 5.

Microfluidic tumor-on-a-chip model for time-resolved shear-stress exposure of S. xylosus–infected tumor cells.

(A) Diagram of the breast cancer–on–a–chip platform used to simulate physiological laminar shear stress. (B) Time-course analysis of tumor-cell viability under laminar shear flow. MDA-MB-231 cells were uninfected or infected with wild-type, ΔdnaK, or ΔdnaK-C Staphylococcus xylosus. Cell viability was quantified at the indicated time points using a live/dead fluorescence assay and normalized to uninfected cells at 0 h. Data represent means ± SEM from three independent microfluidic experiments. (C) Quantification of tumor-cell viability at 4 h of shear-stress exposure. Each data point represents an independent microfluidic experiment. Statistical significance was determined by one-way ANOVA with Tukey’s post hoc test. p < 0.05; ns, not significant.

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