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Fig 1.

Purification of recombinant lumbrokinase.

(A) periplasmic extraction (B) cytoplasmic extraction.

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Table 1.

Characteristics of Collected E. fetida.

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Table 1 Expand

Fig 2.

Map showing the sample collection site with coordinate generated by AutoCAD 2006.

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Fig 3.

Morphological characteristics of locally isolated E. fetida.

(A) Morphology of the collected E. fetida (B) Dissected E. fetida showing internal visceral organ.

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Fig 4.

1% agarose gel electrophoresis.

(A) Lane 2 represents total RNA extracted from earthworm. (B) PCR amplification product with LKF, LKR primers, Lane M; 1Kb DNA ladder, Lane 1: amplified PCR product.

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Fig 5.

Lumbrokinase nucleotide (upper line) obtained from sequencing and deduced amino acid (lower line) sequences.

(GenBank: OP820958; Protein ID: WGV41589).

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Fig 6.

Structural characteristics of lumbrokinase.

(A) The 2D structure prediction lumbrokinase (C represents the coils, H represents the helix and S represents the β-pleated sheets); (B) 3D model of lumbrokinase predicted by I-TASSER (C) Superimposed structure of lumbrokinase and its structural analogue1YM0A.

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Fig 7.

Heterologous expression vector.

(A) recombinant pET22b-lumbrokinase (B) recombinant pET28-SUMO –lumbrokinase.

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Fig 8.

Agarose gel electrophoresis of restriction analysis of recombinant vectors.

(A) pET22b (+) harboring lumbrokinase restricted with EcoR1 and HindIII; (B) pET28a SUMO harboring lumbrokinase restricted with EcoR1 and XhoI. Lane M: 1 kb DNA ladder; Lane 1: Amplified PCR product.

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Fig 9.

SDS-PAGE analysis of total cell proteins from E. coli BL21 (DE3).

(A) harbouring pET22b (+)-lumbrokinase: Lane 1, uninduced sample; Lanes 2-5, samples induced with 1 mM IPTG at 28ºC for 2, 4, 8, and 10 hours, respectively; M, protein molecular weight marker. (B) harbouring pET28a (+) SUMO-lumbrokinase: Lane 1, uninduced sample; Lanes 2-5, samples induced with 1 mM IPTG at 37ºC for 2, 4, 6, and 8 hours, respectively; M, protein molecular weight marker.

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Table 2.

Comparison of activities of lumbrokinase recovered from periplasmic space (expressed in pET-22b) and cytoplasmic space (pET28-a SUMO) from 1liter culture.

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Table 2 Expand

Fig 10.

SDS-PAGE of lumbrokinase at different stages of purification.

(A) expressed with pelB signal: CP, cytoplasmic protein; Un, uninduced sample; P, Periplasmic; (B) expression with SUOMO tag: In, induced sample; Un, un induced sample, H, His taq purified; U, final product after Ulp protease treatment; M, Protein marker.

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Fig 11.

Fibrin plate assay a: Positive control (standard lumbrokinase), b: purified cytoplasmic recombinant lumbrokinase pET-28a (+) SUMO, c: Purified periplasmic recombinant lumbrokinase pET-22b (+), d: Negative control (1 × PBS buffer).

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Fig 12.

Dose-time dependent blood clot lysis: recombinant lumbrokinase and standard lumbrokinase.

1X PBS Buffer negative control (2, 4, and 6 hours of incubation). (A) 0.5 mg/ml standard and recombinant lumbrokinase, tubes 1, 2, and 3 after 2, 4, and 6 hours of incubation, respectively. (B) 2.0 mg/ml standard and recombinant lumbrokinase, tubes 1, 2, and 3 after 2, 4, and 6 hours of incubation respectively.

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Fig 13.

Percentage clot lysis activity: Dose-time dependent blood clot lysis of lumbrokinase.

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Fig 14.

Graph represents the EC50 of recombinant lumbrokinase calculated by AAT Bioquest EC50 calculator.

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