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Fig 1.

Map of the study site, Molokini Shoal Marine Life Conservation District, located in Maui, Hawaii, USA.

Blue triangles show locations of acoustic tracking receivers, yellow and red dots show locations of fish surveys, yellow line shows the boundary of the no-fishing zone inside the crater, and red line shows the boundary of the trolling-only zone around the outside of the crater. Hillshade basemap derived from USGS Digital Elevation Model (DEM) [43]. The partially submerged volcanic crater that forms a wall around the interior is shown in the aerial photo (’Molokini Crater’ by Bossfrog from Wikimedia Commons under the Creative Commons Attribution-Share Alike 4.0 International license).

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Fig 2.

Monthly variation in the number of people visiting Molokini since 2013.

Circles show monthly counts of people at Molokini and are colored by quarters (q) that each corresponds to a 3-month period. The black line is the result of the GAM model. The yellow box denotes the period of travel restrictions associated with the COVID-19 pandemic (from March 2020 to May 2021).

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Fig 3.

Changes in the fish community at Molokini.

Comparisons of fish biomass at all trophic levels among sampling periods from (A) long-term surveys and (C) DAR surveys. Box plots showing median (black line), mean (red line), upper and lower quartiles, and 5th and 95th percentiles. Principal coordinates analysis of fish assemblage composition based on biomass (g m-2) by sampling period for (B) long-term surveys and (D) DAR surveys. Data were ln(x+1)-transformed prior to analyses. Vectors are the relative contribution and direction of influence of fish taxa to the observed variation among sampling periods (Pearson Product movement > 0.5).

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Fig 4.

(A) Fish detections vs. presence of people at Molokini. Grey dots show detections per day for all Caranx melampygus vs. the daily count of people visiting the crater, with darker shade showing higher density. Red line shows GLM fit, grey envelope shows 95% confidence interval. (B) Diel behavior one C. melampygus (44 cm fork length) during 2020 and 2021. Y-axis shows diel cycle with time 00:00 (start of diel cycle) at bottom, noon at center and time 23:59 (end of diel cycle). Horizontal curves show dawn and dusk respectively. Dots show detections of C. melampygus with black representing inside the crater and red outside the crater. Yellow box indicates morning period of high human abundance. An interpretation guide for this figure is provided in the S1 Fig in S1 File.

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