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Fig 1.

Flow diagram of our methods for analyzing wolf scats.

Scats were collected in the field (left), and then processed in three ways: (A) wolf DNA was extracted, genotyped based on microsatellites, and sex was identified; (B) cestode DNA was extracted and genotyped to species; and (C) metabolites of fecal glucocorticoids (molecule shown) were quantified, resulting in a measure of stress. Scat analyses were then matched (right) to known collared wolves, who were monitored throughout their lives, or uncollared wolves. Photo credit: Ellen E. Brandell (wolf scat, left), Jort Vanderveen (collared wolf, right).

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Fig 1 Expand

Table 1.

Fecal prevalence (number of detected infections/number of samples) and apparent maximum prevalence (number of infected wolves/number of unique wolves) in northern Yellowstone wolves years 2018–2020.

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Fig 2.

Proportion of cestode-infected (red) and uninfected (blue) wolves stratified by (A) sex (Female/Male), (B) age class, (C) season, (D) breeding status (0 = non-breeder, 1 = breeder), (E) coat color (Black/Gray), (F) canine distemper virus exposure (Negative/Positive), (G) N. caninum infection (Negative/Positive), and (H) T. gondii infection (Negative/Positive). Stars * and ** denote p < 0.10 and p<0.05 using Fisher’s Exact Test; sample sizes are displayed above columns.

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Fig 2 Expand

Fig 3.

Coefficient estimates for the cestode model (log-odds ratios; points).

Error lines represent 50% (thick) and 95% (thin) confidence intervals. For categorical variables, SEASON winter is with reference to summer, AGE CLASS pup to adult, and SEX male to female.

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Fig 4.

(A) Proportion of scats successfully genotyped when collected within 5 days of deposition (n = 22), or more than 5 days (n = 85), with 95% confidence intervals. Stars ** denote p < 0.05 using Fisher’s Exact Test. (B) Coefficient estimates for the success model (log-odds ratios; points). Error lines represent 50% (thick) and 95% (thin) confidence intervals. Collection PERIOD den and winter are with reference to summer, and COVER is closed with reference to open canopy.

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Fig 4 Expand