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Table 1.

Initial soil physical and chemical properties.

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Table 2.

Total elemental analysis of the amendments.

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Table 2 Expand

Fig 1.

Schematic representation of the experimental setup.

Blue squares represent the wet incubation at 60% WFPS and red-shaded blocks represent the drying period. The samples were collected at the end of days 30 and 72. D, drying phase; W, rewetting phase; WFPS, water-filled pore spaces. D-W1, 30 d 60% WFPS; D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d; D-W3, 72 d 60% WFPS. * Indicates sampling points.

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Table 3.

Reagents used for extracellular enzymes assay.

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Table 3 Expand

Fig 2.

The interaction of soil type × moisture cycle × amendment for the potential β-glucosidase, β = glucosaminidase and acid phosphatase (panels a, b and c, respectively) in the amended soil mesocosms. Asterisk indicates difference among treatments (α = 0.05), based on three-way ANOVA. Error bars represent standard errors (n = 3). C, control; HR, hemp; HB, hemp biochar; HA, hardwood biochar; D-W1, 30 d 60% WFPS; D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d; D-W3, 72 d 60% WFPS.

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Fig 3.

The interaction of soil type × moisture cycle × amendment for the potential arylsulfatase, phosphodiesterase, and geometric mean of enzyme activities (panels a, b and c, respectively) in the amended soil mesocosms. Asterisk indicates difference among treatments (α = 0.05), based on three-way ANOVA. Error bars represent standard errors (n = 3). C, control; HR, hemp; HB, hemp biochar; HA, hardwood biochar; D-W1, 30 d 60% WFPS; D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d; D-W3, 72 d 60% WFPS.

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Fig 3 Expand

Fig 4.

The interaction of soil type × moisture cycle × amendment for the total PFLA bacteria biomass in the amended soil mesocosms.

Asterisk indicates difference among treatments (α = 0.05), based on three-way ANOVA. Error bars represent standard errors (n = 3). C, control; HR, hemp; HB, hemp biochar; HA, hardwood biochar; D-W1, 30 d 60% WFPS; D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d; D-W3, 72 d 60% WFPS.

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Fig 4 Expand

Table 4.

Means (n = 3) for total PLFA biomass, total PLFA fungi biomass, fungi:bacteria ratio, G (+): G(-), and sat:unsat for interaction effects (soil type × day × amendment) in soil mesocosms.

Lower case letters designate LSMeans comparisons. All statistics are presented when treatment effects were significant (α = 0.05).

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Fig 5.

The interaction of soil type × moisture cycle × amendment for the pH, nitrate nitrogen, permanganate oxidizable carbon, and total carbon (panels a, b, c, and d, respectively) in the amended soil mesocosms. Asterisk indicates difference among treatments (α = 0.05), based on three-way ANOVA. Error bars represent standard errors (n = 3). C, control; HR, hemp; HB, hemp biochar; HA, hardwood biochar; D-W1, 30 d 60% WFPS; D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d; D-W3, 72 d 60% WFPS.

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Fig 6.

Principal components analysis (panel a: plot of eigen plot of the eigenvectors, panel b: plot of the PC scores) of soil enzymes and phospholipid-derived fatty acid (PLFA) profiles as impacted by amendments and moisture cycles in the Coastal soil. D-W1, 30 d 60% WFPS; D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d; and D-W3, 72 d 60% WFPS. C, control; HR, hemp residue; HB, hemp biochar; HA, hardwood biochar. GEOM, geometric mean of enzyme activities; POXC, permanganate oxidizable carbon; TC, total soil organic carbon; TN, total nitrogen; GramPost_GramNeg, gram-positive to gram-negative bacteria biomass; Sat_Unsat, ratio of saturated to unsaturated fatty acids.

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Fig 7.

Principal components analysis (panel a: plot of eigen plot of the eigenvectors, panel b: plot of the PC scores) of soil enzymes and phospholipid-derived fatty acid (PLFA) profiles as impacted by amendments and moisture cycles in the Piedmont soil. D-W1, 30 d 60% WFPS; D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d; and D-W3, 72 d 60% WFPS. C, control; HR, hemp residue; HB, hemp biochar; HA, hardwood biochar. GEOM, geometric mean of enzyme activities; POXC, permanganate oxidizable carbon; TC, total soil organic carbon; TN, total nitrogen; GramPost_GramNeg, gram-positive to gram-negative bacteria biomass; Sat_Unsat, ratio of saturated to unsaturated fatty acids.

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Fig 8.

Pearson correlation matrix of soil enzymes, microbial community structure and biochemical properties of soil in Coastal Plain soil in D-W1, 30 d 60% WFPS (panel a); D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d (panel b); D-W3, 72 d 60% WFPS (panel c). GEOM, geometric mean of enzyme activities; POXC, permanganate oxidizable carbon; TC, total soil organic carbon; TN, total nitrogen; GramPost_GramNeg, gram-positive to gram-negative bacteria biomass; Sat:Unsat, ratio of saturated to unsaturated fatty acids. *p<0.05, **p<0.01, ***p<0.0001.

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Fig 9.

Pearson correlation matrix of soil enzymes, microbial community structure and biochemical properties of soil in Piedmont soil in D-W1, 30 d 60% WFPS (panel a); D-W2, 30 d 60% WFPS followed by 7 d alternate D-W for 42 d (panel b); D-W3, 72 d 60% WFPS (panel c). GEOM, geometric mean of enzyme activities; POXC, permanganate oxidizable carbon; TC, total soil organic carbon; TN, total nitrogen; GramPost_GramNeg, gram-positive to gram-negative bacteria biomass; Sat:Unsat, ratio of saturated to unsaturated fatty acids. *p<0.05, **p<0.01, ***p<0.0001.

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Fig 9 Expand