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Fig 1.

Amino acid sequence of the PHF-Tau protein with the observed b-strand regions and the corresponding schematic view of the C-shaped architecture of the protofilament core with possible high-affinity binding sites of the protofibril (S1 to S5) [35,36].

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Fig 1 Expand

Fig 2.

Molecular structures and its 3D geometric representations of (a) 3-alpha-cholesterol, (b) 1-palmitoyl-2-oleoyl-sn-glycero-3-phosphocholine, and (c) C18:1 sphingomyelin.

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Fig 2 Expand

Fig 3.

(a) The docking conformation of 3-alpha-cholesterol on PHF-Tau showing the H-bond formation between the PHF-Tau backbone and OH group of the ligand. (b) Electrostatic surface potential model on the docking site of 3-alpha-cholesterol on PHF-Tau which shows the hydrophobicity of the binding site (blue regions are hydrophilic and red orange regions are hydrophobic). (c) H-bonds obtained in NBO analysis.

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Fig 3 Expand

Fig 4.

(a) The docking conformation of 1-POPC on PHF-Tau showing the H-bond formation between 1-POPC and the PHF-Tau backbone. (b) Electrostatic surface potential model on the docking site of 1-POPC ON PHF-Tau which shows the hydrophobicity of the binding site (blue regions are hydrophilic and red orange regions are hydrophobic).

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Fig 4 Expand

Fig 5.

Intermolecular interactions between 1-POPC and PHF-Tau based on NBO analysis.

Interatomic distance is shown in angstrom.

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Fig 5 Expand

Fig 6.

(a) The docking conformation of C18:1 Sphingomyelin on PHF-Tau showing the H-bond formation between C18:1 Sphingomyelin and the PHF-Tau backbone. (b) Hydrophobicity of the binding site (blue regions are hydrophilic and red orange regions are hydrophobic).

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Fig 6 Expand

Fig 7.

NBO analysis of the intermolecular interactions between PHF-Tau and C18:1 Sphingomyelin.

(a) H-bond formation between C18:1 Sphingomyelin and the PHF-Tau backbone and (b) significant London dispersion forces. Interatomic distances are shown in angstrom.

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Fig 7 Expand

Fig 8.

Variations of RMSD values in molecular dynamics simulations of the three membrane lipids.

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Fig 8 Expand

Fig 9.

Kinetic, potential and temperature variations in molecular dynamics simulations of of 3-alpha-cholesterol docked structure with RMSD = 0.

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Fig 9 Expand

Fig 10.

Kinetic, potential and temperature variations in molecular dynamics simulations of 1-POPC docked structure with RMSD = 0.

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Fig 10 Expand

Fig 11.

Kinetic, potential and temperature variations in molecular dynamics simulations of C18:1Sphingomelin docked structure with RMSD = 0.

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Fig 11 Expand

Fig 12.

Variations of RMSD values in molecular dynamics simulation of PHF-Tau protofilament and PHF-Tau protofilament with 1-POPC.

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Fig 12 Expand

Fig 13.

Ligand (AM1-BCC optimized structure) embedded in PHF-Tau protofilament (tail side); RMSD = 0.

Hydrophobicity surface is shown. Solvent used is water. Inset shows the preferred configuration of the the ligand: (a) cholesterol; (b) 1-palmitoyl-2-oleoyl-sn-glycero-3-phosphocholine; and (c) C18:1 sphingomyelin and their interactions with the protein.

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Fig 13 Expand

Fig 14.

The docked structures of lipids with RMSD = 0 in solvated and unsolvated PHF-Tau.

(a) cholesterol; (b) 1-palmitoyl-2-oleoyl-sn-glycero-3-phosphocholine; and (c) C18:1 sphingomyelin.

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Fig 14 Expand

Fig 15.

Variations of RMSD values in molecular dynamics simulations of the three membrane lipids in solvated protein.

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Fig 15 Expand