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Fig 1.

Synthetic route for hydrazone derivatives H1-H5.

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Fig 2.

(A) Effect of H1 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (B) Effect of H2 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.), and indomethacin (20 mg/kg, i.p.); (C) Effect of H3 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (D) Effect of H4 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (E) Effect of H5 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.) in the acetic acid-induced writhing test in mice (n = 6, per group). Values are expressed as the mean ± SEM, where a indicates p < 0.05, significantly different from the control group, according to ANOVA, followed by Tukey’s test.

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Fig 3.

(A) Effect of H1 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (B) Effect of H2 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (C) Effect of H3 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (D) Effect of H4 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (E) Effect of H5 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.) in the first phase of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± SEM, where a indicates p < 0.05, significantly different from the control group, followed by Tukey’s test.

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Fig 3 Expand

Fig 4.

(A) Effect of H1 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (B) Effect of H2 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (C) Effect of H3 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (D) Effect of H4 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.); (E) Effect of H5 (20 and 40 mg/kg, p.o.), morphine (10 mg/kg, i.p.) and indomethacin (20 mg/kg, i.p.) in the second phase of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± SEM, where a indicates p < 0.05, significantly different from the control group, b indicates p < 0.05 in comparison with indomethacin group and c indicates p<0.05 in comparison with morphine group, according to ANOVA, followed by Tukey’s test.

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Fig 4 Expand

Fig 5.

Effect of H5 (20 mg/kg, p.o.), naloxone (1.5 mg/kg, i.p.), naloxone + H5 and morphine (10 mg/kg, i.p.) in the first (A) and second (B) phases of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± S.E.M., where a indicates p < 0.05 in comparison with control group, b indicates p<0.05 in comparison with morphine group, and c indicates p < 0.05 in comparison with H5 group, according to ANOVA, followed by Tukey’s post-test.

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Fig 6.

Effect of L-NAME (20 mg/kg, i.p.), H5 (20 mg/kg, p.o.), L-NAME + H5, in the first (A) and second (B) phases of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± S.E.M., where a indicates p < 0.05 in comparison with control group, according to ANOVA, followed by Tukey’s post-test.

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Fig 6 Expand

Fig 7.

Effect of ondansetron (0.5 mg/kg, i.p.), H5 (20 mg/kg, p.o.), ondansetron + H5 in the first (A) and second (B) phases of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± S.E.M., where a indicates p < 0.05 in comparison with control group and b indicates p < 0.05 in comparison with the ondansetron group, according to ANOVA, followed by Tukey’s post-test.

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Fig 8.

Effect of atropine (0.1 mg/kg, i.p.), H5 (20 mg/kg, p.o.), atropine + H5 in the first (A) and second (B) phases of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± S.E.M., where a indicates p < 0.05 in comparison with control group, according to ANOVA, followed by Tukey’s post-test.

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Fig 9.

Effect of glibenclamide (2 mg/kg, i.p.), H5 (20 mg/kg, p.o.), glibenclamide + H5 in the first (A) and second (B) phases of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± S.E.M., where a indicates p < 0.05 in comparison with control group, according to ANOVA, followed by Tukey’s post-test.

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Fig 9 Expand

Fig 10.

Effect of ruthenium red (3 mg/kg, i.p.), H5 (20 mg/kg, p.o.), ruthenium red + H5 in the first (A) and second (B) phases of the formalin-induced nociception test in mice (n = 6, per group). Values are expressed as the mean ± S.E.M., where a indicates p < 0.05 in comparison with control group, according to ANOVA, followed by Tukey’s post-test.

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Fig 11.

Effect of H5 (20 and 40 mg/kg, p.o.) and dexamethasone (2 mg/kg, i.p.) on leukocyte migration into the peritoneal cavity induced by carrageenan in mice.

Values are expressed as the mean ± S.E.M. (n = 6, per group), where a indicates p < 0.05, significantly different from the control group, according to ANOVA, followed by Tukey’s test.

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Fig 12.

Effect of H5 (20 and 40 mg/kg, p.o.) and indomethacin (20 mg/kg, i.p.) on paw edema induced by carrageenan in mice.

The sham group was treated only with saline, whereas the control group received saline and carrageenan. Values are expressed as the mean ± S.E.M. (n = 6, per group), where a indicates p < 0.05, significantly different from the control group, according to ANOVA, followed by Tukey’s post-test.

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Fig 13.

Effect of H5 (20 mg/kg, p.o.) on paw edema induced by histamine in mice.

The sham group was treated only with saline, whereas the control group received saline and histamine. Values are expressed as the mean ± S.E.M. (n = 6, per group), where a indicates p < 0.05, significantly different from the control group, according to ANOVA, followed by Tukey’s post-test.

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Fig 14.

Effects of H5 (20 and 40 mg/kg, p.o.), and diazepam (2.5 mg/kg, i.p.) in the Rota-rod test in mice.

Values are expressed as mean ± S.E.M. (n = 6, per group), where a indicates p < 0.05, significantly different from the control group, according to ANOVA, followed by Tukey’s post-test.

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Fig 14 Expand

Table 1.

Lethality rate of Artemia salina nauplius to hydrazone (H5).

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Table 2.

Artemia salina toxicity test of hydrazone H5.

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Fig 15.

Results of the docking procedures for H5 and Meloxicam, (A) Interaction profile of H5 in the murine COX-2 enzyme binding site after the docking study; (B) Interaction profile Meloxicam in the murine COX-2 enzyme binding site after the redocking study.

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Table 3.

Comparative in silico physicochemical properties and ADMET profile of the anti-inflammatory drugs indomethacin and meloxicam and new hydrazone series H1 to H5.

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Table 3 Expand