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Fig 1.

(A) The structures of the S protein trimer (electrostatic potential surface area (PDB ID: 6vxx)),S1(A), S2(A), S1(B), S2(B), S1(C) and S2(C) are shown in cyan, gray, blue, orange, magenta, and yellow-green, respectively. (B) Schematic of the SARS-CoV-2 S protein primary structure, colored by domain.

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Fig 1 Expand

Table 1.

The highest binding energy of drug molecules with the S protein in molecular docking simulations.

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Table 1 Expand

Fig 2.

(A) Binding mode of the interaction of tizoxanide with the S protein. The S protein trimer is shown as a transparent red surface, S1(A), S1(B), and S2(C) are shown as cyan, blue, and yellow-green cartoons, respectively, and tizoxanide is shown as green spheres. (B) Binding mode of tizoxanide with the S protein. S1(A), S1(B), and S2(C) are shown as cyan, blue, and yellow-green surfaces, respectively, and tizoxanide is shown as green sticks. (C) The key residues that may form potential interactions with tizoxanide (green sticks) and polar interactions are indicated by blue dotted lines.

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Fig 2 Expand

Fig 3.

(A) Binding mode of the interactions of bictegravir and dolutegravir with the S protein. The S protein trimer is shown as a transparent red surface, S1(A), S1(B), and S2(B) are shown as cyan, blue, and orange cartoons, respectively, and bictegravir and dolutegravir are shown as green spheres. (B) and (D) Binding mode of bictegravir and dolutegravir (green) with the S protein. S1(A) and S1(B) are shown as cyan and blue surfaces, and bictegravir and dolutegravir are shown as green sticks. (C) and (E) The key residues that may form potential interactions with bictegravir and dolutegravir (green sticks) and polar interactions are indicated by blue dotted lines.

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Fig 3 Expand

Fig 4.

(A) Binding mode of the interaction of arbidol with the S protein. The S protein trimer is shown as a transparent red surface; S1(A), S2(A), and S2(B) are shown as cyan cartoons; and arbidol is shown as a green sphere. (B) Binding mode of arbidol (green) with the S protein. S1(A), S2(A), and S2(B) are shown as cyan surfaces, and arbidol is shown as a green stick. (C) The key residues that may form potential interactions with arbidol (green stick) and polar interactions are indicated by blue dotted lines.

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Fig 4 Expand

Fig 5.

(A) and (C) The amino acid residues that surround the phenol group and nitro group of tizoxanide (green). (B) and (D) The amino acid residues surrounding the phenol and nitro groups are shown as green surfaces.

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Fig 5 Expand

Fig 6.

Strategy for further structural optimization of tizoxanide.

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Fig 6 Expand

Fig 7.

Molecular mode results for Ti-1(A), Ti-2(B), and Ti-3(C) with the S protein and the key residues that may form potential interactions with compounds Ti-1, Ti-2 and Ti-3.

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Fig 7 Expand

Fig 8.

(A) The amino acid residues that surround the trifluorophenyl and difluorophenyl moieties of bictegravir (green) and dolutegravir (blue-green). (B) The amino acid residues surrounding the trifluorophenyl and difluorophenyl moieties are shown as green surfaces.

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Fig 8 Expand

Fig 9.

Strategy for further structural optimization of bictegravir and dolutegravir.

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Fig 9 Expand

Fig 10.

Molecular mode results for BD-1(A), BD-2(B), and BD-3(C) with the S protein and the key residues that may form potential interactions with compounds BD-1, BD-2, and BD-3.

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Fig 11.

(A) The polar residues that surround the Br of arbidol (green). (B) The H-binding region consists of polar residues, which are shown as a transparent surface. (C) The hydrophobic residues that surround the N-methyl group. (D) The arbidol–hydrophobic residue complex; the hydrophobic residues are shown as a transparent surface.

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Fig 11 Expand

Fig 12.

Strategy for further structural optimization of arbidol.

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Fig 12 Expand

Fig 13.

Molecular mode results for Ar-1 (A), Ar-2 (B), and Ar-3 (C) with the S protein and the key residues that may form potential interactions with compounds Ar-1, Ar-2, and Ar-3.

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Fig 13 Expand