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Table 1.

Voucher numbers, localities, and GenBank accession numbers for genetic samples.

DRC = Democratic Republic of the Congo; RC = Republic of Congo; SA = South Africa; GNP = herpetological collection of the E. O. Wilson Biodiversity Center, Gorongosa National Park, Mozambique. Other collection acronyms are explained in Sabaj [108]. Note that Lawson et al. [109] erroneously listed the specimen of Atractaspis sp. as MVZ 228653.

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Table 1 Expand

Fig 1.

Map of sub-Saharan Africa and western Asia/Middle East, showing sampling localities for atractaspidines used in this study.

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Fig 1 Expand

Fig 2.

Maximum-likelihood phylogeny of Atractaspidinae with combined 16S, ND4, cyt b, c-mos, and RAG1 data sets.

Closed circles denote clades with Bayesian posterior probability values ≥ 0.95. Diamonds denote clades with strong support in both maximum likelihood analyses (values ≥ 70) and Bayesian analyses (posterior probability values ≥ 0.95).

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Fig 2 Expand

Table 2.

Primers used for sequencing mitochondrial and nuclear genes.

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Table 2 Expand

Fig 3.

Maximum-likelihood phylogeny of Atractaspidinae and Aparallactinae with combined 16S, ND4, cyt b, c-mos, and RAG1 data sets.

Diamonds denote clades with maximum likelihood values ≥ 70 and Bayesian posterior probability values ≥ 0.95; closed circles denote clades with Bayesian posterior probability values ≥ 0.95.

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Fig 3 Expand

Fig 4.

Phylogeny resulting from BEAST, based on four calibration points.

Nodes with high support (posterior probability ≥ 0.95) are denoted by black circles. Median age estimates are provided along with error bars representing the 95% highest posterior densities (HPD) (Table 3).

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Fig 4 Expand

Table 3.

Estimated dates and 95% highest posterior densities (HPD) of main nodes.

Node labels correspond to those in Fig 4.

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Table 3 Expand

Fig 5.

Computed tomography (CT) scans of aparallactine and atractaspidine genera.

Homoroselaps lacteus (CAS 173258) (A); Atractaspis bibronii (CAS 111670) (B); Chilorhinophis gerardi (CAS 159106) (C); Polemon christyi (CAS 147905) (D); Aparallactus niger (AMNH 142406) (E); Aparallactus modestus (CAS 111865) (F); Aparallactus capensis (G); Macrelaps microlepidotus (H); Amblyodipsas polylepis (CAS 173555) (I); Xenocalamus bicolor (CAS 248601) (J).

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Fig 5 Expand

Fig 6.

Ancestral-state reconstructions with ML optimization on the ML trees from the concatenated analyses shown in Fig 2.

(A) fang morphology, (B) dietary preference. Aparallactus 1 = A. niger; Aparallactus 2 = A. modestus; Aparallactus 3 = A. capensis, A. cf. capensis, A. guentheri, A. jacksonii, A. lunulatus, and A. werneri; Amblyodipsas 1 = A. concolor; Amblyodipsas 2 = A. dimidiata, A. polylepis, and A. unicolor; Amblyodipsas 3 = A. ventrimaculata; Amblyodipsas 4 = A. microphthalma.

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Fig 6 Expand