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Fig 1.

Graphical abstract.

Conceptual diagram describing the three-steps approach of the present work (graphical abstract).

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Fig 2.

Location of the sampling stations.

Location and sampling design of the empirical data sets used in the current work, bottom trawl survey (EVHOE 2015, left panel), small pelagic surveys using acoustic techniques (PELGAS 2015; middle panel) and worldwide reference of the areas (right panel).

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Table 1.

Summary of the indicators used in the current work, organized by group.

Underlined indices are the ones retained for other analysis.

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Fig 3.

Relationships among spatial indicators.

Correlation matrix plot of the indicators estimated for each annual species distribution, and respective cluster groups (on top) that represent the three categories (colored accordingly).

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Fig 4.

Species groups according to the indicators.

The results of each indicator for all years, were averaged by species and scaled. Left panel represents the heatmap with respective cluster (groups defined by kmeans) and right panel, shows the PCA results. Indicators and species names/symbols are colored according to the categories or cluster groups, respectively.

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Fig 5.

Relationships between quantity-occupancy-aggregation for all species under study.

Spearman correlation between the three aspects of species spatial behavior (years pooled): Aggregation (measured by Gini index), Occupancy (measured by the percentage of presence) and Quantity (measured by the mean log biomass). The width of the edge line represents the strength of the relationship, its color the direction (positive in blue and negative in red) and the value in the middle, is the correlation coefficient. Non-significant correlations were omitted. Codes for the species can be found in the text.

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Fig 6.

Groups of species with similar quantity-occupancy-aggregation relationships.

See further details in the text.

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