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Table 1.

Insulin glargines employed in this study.

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Fig 1.

Insulin glargine (Glaritus) lots 32 and 96 induce heightened innate activity in the MIMIC® PTE.

MIMIC® PTE cultures were treated with different batches of insulin glargines at 30 nM (5 U/ml) for 48 hours. Thereafter, the culture supernatant were collected and evaluated for the secretion of different cytokines by multiplex assay. The plotted values represent mean ± SEM (pg/ml) of IL-8 secretion (A) and IL-6 secretion (B) for 12–14 donors. (C) Graphical representation of Pearson correlation analysis using confidence intervals (CI) of IL-8 and IL-6 secretion induced by all insulin glargine products. **, p<0.01; ****, p<0.001; -, No treatment; L+R, LPS+R848; Bas, Basalin; B, Bonglixan; HI, Human Insulin (Insuman). Lot numbers represent the last two digits of the lot numbers shown in Table 1.

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Table 2.

Statistical analysis of IL-8 and IL-6 secretion: Glaritus versus Lantus.

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Fig 2.

Removal of insulin glargine from Glaritus lots 32 and 96 abrogates innate immune activity induced by these formulations.

Insulin glargines were incubated overnight with the L6B10 anti-insulin mAb and then processed through a 30 kDa filter. (A) Pre- and post-filtration samples were analyzed by a 12% reducing SDS-PAGE gel; the insulin glargine band is observed at 6–10 kDa. (B) Pre- and post-filtration samples were analyzed by 48-hour MIMIC® PTE assay, then multiplex assay for IL-8 production. Data from 19 healthy donors were plotted as % IL-8 secretion over no-treatment control. (C) MIMIC® PTE cultures were treated with preservative and non-preservative insulins at doses of 30, 50, and 500 nM and then examined for IL-8, IL-6, and MCP-1 secretion by multiplex assay. Data is plotted as mean ± SEM (pg/ml) from 8–13 healthy donors. ****, p <0.001.

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Fig 3.

The innate response induced by insulin glargine in MIMIC® PTE assays is largely driven by insulin signaling through the IRA/IRB and MEK pathways.

MIMIC® PTE cultures were incubated with the indicated treatments immediately following PBMC application. (S961 is the insulin receptor AB antagonistic peptide.) 1 hour later, 30 nM (5 U/ml) insulin glargines were added to the wells and incubated for 48 hours. Thereafter, the culture supernatants was collected and analyzed for IL-8 secretion by multiplex assay. Data represented as mean ± SEM of 8–12 healthy donors. Lot numbers represent the last two digits of the lot numbers shown in Table 1.

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Fig 4.

Glaritus lots 32 and 96 show differential by-product profiles by SEC-HPLC analysis.

(A) 500 ng insulin glargines were analyzed by 3–12% gradient Bis-Tris Native-PAGE under non-reducing conditions followed by silver staining. Molecular markers were used as size standards. Images were taken with a Kodak GL 1500 Imaging system and the insulin glargine band was observed at 6–10 kDa. (B) Representative SEC-HPLC analysis of Glaritus lots 32 and 96 compared with the originator lot 06. Arrows indicate the presence of distinct peaks observed between the originator and non-originator insulin glargines.

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