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Fig 1.

Illustration of a nest and a maze of complexity levels (a) 0, (b) 1, (c) 2 and (d) 3. Each circle represents a passage and a decision to make (light gray ones are correct decision, leading to the food reward, and dark grey ones are wrong decisions). The larger circle on the right upper (a, c) side or right lower side (b, d) represents the food reward (honey). The correct number of decisions, including the decision to enter the maze are 1, 2, 3, and 4 for complexity levels 0 to 3, respectively. The total number of decisions are 1, 3, 7, and 15 for the same complexity levels. Mazes included either right or left turns as the correct ones, which were randomly applied per colony.

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Fig 1 Expand

Table 1.

The effect of colony size and maze complexity on the three foraging response variables on the first day and test.

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Table 1 Expand

Fig 2.

(a) The interaction between colony size and maze complexity in their effect on food-discovery time on the first test; (b) the decrease in food-discovery time with successive tests performed on the same day. Letters denote significant differences according to a post-hoc comparison.

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Fig 2 Expand

Table 2.

The effect of colony size, maze complexity and day (1 vs. 2) on food-discovery time, the number of workers searching, and food consumption (N = 19 for food-discovery time and workers searching and 17 for food consumption).

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Table 2 Expand

Fig 3.

(a) The decrease in food-discovery time between the first tests on two successive days; (b) the positive correlation between colony size and the number of workers searching (mean values for the first tests on the two successive days are presented). The asterisk indicates a significant difference.

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Fig 3 Expand

Table 3.

The effect of day (before and after the time interval) and the time interval between days on the three foraging response variables.

N = sample size (number of colonies).

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Table 3 Expand

Fig 4.

The effect of the time interval (in days, horizontal axis) between tests on the difference between food-discovery time before and after this interval (Δ discovery time, vertical axis).

Values above zero indicate that colonies reached the food reward faster after the time interval than before it, while values below zero indicate that colonies reached the food reward faster before the time interval. As intervals become large, food-discovery time returns to the basic initial levels (around zero), which takes place according to this model after ~16 days.

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Fig 4 Expand