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Fig 1.

Morphologically diverse marine microbes.

Length-to-width ratio versus surface-equivalent sphere diameter (ESD) [13] of some abundant free-living microbes in the upper ocean (S1 Table). Surface area (S) of ESD () was calculated based on the assumption of a prolate spheroid [13]. Color of points shows average abundances. Dashed black line represents the length: width ratio of a perfectly spherical cell.

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Fig 2.

Grazing by Oikopleura dioica influences particle fate.

Different fates of particles grazed by the appendicularian Oikopleura dioica: particles associated with the discarded house (white arrow) via retention on either the inlet filters (IF), food-concentrating filter (FCF), or house walls; particles captured on the pharyngeal filter (PF), ingested, and incorporated into fecal pellets (FP) (blue arrow). Arrow widths represent the average flux of houses (703 mg C m-2 d-1) and fecal pellets (446 mg C m-2 d-1) [44]. Arrow lengths represent the average sinking rates of houses (50 m day-1) ([38] and references therein) and fecal pellets (60 m day-1) [44]. Values show the range of flux for houses ([38] and references therein) and fecal pellets [42, 44], and the sinking rates of houses [42] and fecal pellets [44, 45]. Schematic of O. dioica by Jenna Valley.

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Fig 3.

Experimental bead mixture for incubations.

(A) Schematic of experimental bead mixture used in each of three incubation experiments with Oikopleura dioica. (B) Top: O. dioica in house filtering a mixture of Rhinomonas reticulata (red, ~17 μm diameter) and fluorescent 10 μm microspheres (green). PF: pharyngeal filter; FCF: food-concentrating filter. Scale bar 0.5 mm. Bottom: experimental bead mixture (3–10 μm) in the gut post-incubation. Scale bar 50 μm.

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Table 1.

Experimental conditions for three incubation experiments.

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Table 2.

Observational results from incubation experiments with Oikopleura dioica.

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Fig 4.

Fate of different shaped beads from three incubation experiments.

Relative proportions (mean ± SD) of various bead mixes in the ambient water at the start of the incubation (T0), gut, and house of the appendicularian Oikopleura dioica.

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Fig 5.

Particle shape affects selection by the appendicularian Oikopleura dioica.

Selectivity coefficients (mean Chesson’s α-index ± SE) for different bead types in the houses and guts from each of three incubation experiments with Oikopleura dioica. * indicates selectivity values that were significantly different from non-selectivity (α = 0.33, dashed line) tested using t-tests with a Bonnferoni correction of alpha level (p ≤ 0.0028) (Table 3).

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Table 3.

Statistical results for incubation experiments.

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Fig 6.

Trajectories and orientation of spheroidal microbeads through the feeding-filter of Oikopleura dioica.

(A) Frequency histograms show the angles for prolate spheroid microbeads (7.8 x 22 μm) suspended in the fluid of the food-concentrating filter and adhered to the filter mesh of Oikopleura dioica. Sectors correspond to number of beads observed, Ф is the grand mean direction for all beads, N is the number of independently measured beads, n is the instantaneous angles pooled for all beads. Angle measurements for beads adhered to the mesh represent independent measurements for N = 40 beads, whereas measurements for beads suspended in the fluid represent instantaneous angles pooled for all individuals, n = 192, from the trajectories of N = 10 beads. All angles are relative to the fluid flow. (B) Five sample trajectories of beads transported through the food-concentrating filter (S1 Video). Frames were colored-coded (red, yellow, green, cyan, blue, magenta) so that the color order shows direction and white indicates a particle has not moved. Arrows show directions of fluid flow. Montages show the bead orientations for the respective trajectory.

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