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Fig 1.

Experimental setup.

(A) Individual client fish (Indo-Pacific Naso elegans) is introduced to a novel object (a white ball), (B) Individual client fish is introduced to an allopatric cleaner species (the Caribbean cleaning goby Elacatinus evelynae), (C) Individual client fish is introduced to a sympatric cleaner species (the Indo-Pacific cleaner wrasse Labroides dimidiatus) and (D) Individual client fish is introduced to a conspecific. Images of fish are not in real scale, particularly those of the cleaner fish L. dimidiatus and E. evelynae when compared with client N. elegans size.

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Fig 1 Expand

Table 1.

Frequency of observed behavioural measures for each experimental treatment.

These include: a) the frequency of cleaning interactions, b) mean inspection duration, c) proportion of interactions in which tactile stimulation was applied to clients, d) frequency of jolts per 100 s of inspection, e) frequency of chases, f) duration of chases and g) frequency of bites given by focal individual. Mean ± Standard Error (SEM) are provided for each behavioural measure.

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Fig 2.

Whole brain nonapeptide’ levels in clients (Naso elegans).

(A) levels of arginine vasotocin (AVT) and (B) levels of isotocin (IT) in four treatment groups: a) sympatric cleaner (Labroides dimidiatus), b) allopatric cleaner (Elacatinus evelynae), c) conspecific (N. elegans) and d) ball, expressed as AVT (pmol/mg) and IT (pmol/mg). Medians (full lines) and interquartile range are presented in boxes.

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Fig 2 Expand

Fig 3.

Levels of arginine vasotocin (AVT) in different brain macro-areas: Forebrain, diencephalon, optic tectum, cerebellum and brain stem in clients (Naso elegans).

Four treatment groups: a) sympatric cleaner (Labroides dimidiatus), b) allopatric cleaner (Elacatinus evelynae), c) conspecific (N. elegans) and d) ball, expressed as AVT (pmol/mg). Medians (full lines) and interquartile range are presented in boxes.

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Fig 3 Expand

Fig 4.

Levels of isotocin (IT) in different brain macro-areas: Forebrain, diencephalon, optic tectum, cerebellum and brain stem in clients (Naso elegans).

Four treatment groups: a) sympatric cleaner (Labroides dimidiatus), b) allopatric cleaner (Elacatinus evelynae), c) conspecific (N. elegans) and d) ball, expressed as IT (pmol/mg). Medians (full lines) and interquartile range are presented in boxes.

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Fig 4 Expand