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Table 1.

Opportunistic observations of actively walking and non-walking epibenthic hard clams, Mercenaria mercenaria, made while diving in the Peconic Bays, New York, USA, June 2007 –June 2016.

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Table 2.

Summary of factors hypothesized to affect specific metrics of observed hard clam walking behavior in the Peconic Bays, New York, USA.

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Table 3.

Summary of factors hypothesized to differ between adjacent (~50–60 m apart) areas at the East Marion site where hard clam walking was frequently observed (‘walking area’) versus rarely observed (‘non-walking’ area).

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Fig 1.

Hard clam, Mercenaria mercenaria, walking at East Marion, New York, USA.

(A) Active walker illustrating typical shell orientation: ligament parallel to sediment surface, siphon above sediment in furrow. (B) Active walker with freshly excavated trail (furrow). (C) Non-walking clam lying on sediment surface, next to active walker. (D) Active walker at end of semi-circular trail. (E) Epibenthic clams, with posterior ends (where siphons are located) adjacent to one another.

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Fig 2.

Bar plots of mean (±1 standard error of the mean, SE) Mercenaria mercenaria density versus site/walking designation in Orient Harbor, New York, USA.

(A) Total (epibenthic + endobenthic) clam density. (B) Surface (epibenthic) clam density. All densities quantified at the East Marion site, except where noted as * (= Peter’s Neck site). W = walking clams, in ‘walking’ areas; NW = non-walking clams, in ‘non-walking’ area; WNW = non-walking clams in ‘walking’ area.

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Fig 3.

Morisita’s Index for Mercenaria mercenaria versus site/walking designation in Orient Harbor, New York, USA.

W = walking clams, in ‘walking’ areas; NW = non-walking clams, in ‘non-walking’ area; WNW = non-walking clams in ‘walking’ area. Dashed line demarcates transition from a random (IM = 1) to an aggregated distribution (IM > 1); asterisk denotes a significant χ2 test: p = 0.001.

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Fig 4.

Bar plots of mean (±1 standard error of the mean, SE) nearest neighbor distances of Mercenaria mercenaria at East Marion, New York, USA.

(A) endo- + epibenthic clams and (B) epibenthic clams only. W = walking clams, in ‘walking’ area; NW = non-walking clams, in ‘non-walking’ area; WNW = non-walking clams in ‘walking’ area. Different letters above bars represent significantly different densities at p < 0.05.

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