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Table 1.

Genes and primer sequences of Calanus sinicus used in this study.

GAPDH, EF1ɑ and 16S are housekeeping genes, while the other six are target genes.

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Table 1 Expand

Fig 1.

Sampling station for Calanus sinicus.

The sampling station is marked with a star (☆) and is located inside the Yellow Sea Cold Water Mass (YSCWM). The boundary of the YSCWM is defined by the 10°C bottom isotherm and marked with bold black line.

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Fig 1 Expand

Fig 2.

Vertical profiles of temperature and Chl a concentration at the Calanus sinicus sampling station.

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Fig 2 Expand

Fig 3.

Oxygen consumption rate (OCR) of Calanus sinicus cultured under different temperatures by food environments.

9NF = 9°C not fed, 9F = 9°C fed, 18NF = 18°C not fed and 18F = 18°C fed.

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Fig 4.

The composition time series of developmental stages and jaw phases in Calanus sinicus.

(a) 9NF (9°C not fed), (b) 18NF (18°C not fed), (c) 9F (9°C fed) and (d) 18F (18°C fed).

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Fig 4 Expand

Fig 5.

Jaw phase morphology of Calanus sinicus.

(a) pre-apolysis on Day 0; (b) post-apolysis on Day 6 in Group 9F (9°C fed), tooth forming; (c) post-apolysis on Day 9 in Group 9F. Scale bars: 20 μm.

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Fig 5 Expand

Fig 6.

Relative gene expression variation of genes associated with the molting process of Calanus sinicus.

(a) EcR (ecdysteroid receptor) expression. (b) FAMeT (Farnesoic acid O methyltransferase) expression. 9NF = 9°C not fed, 9F = 9°C fed, 18NF = 18°C not fed and 18F = 18°C fed. D0 to D16 represent Days 0 to 16. The expression level was normalized to the geometric mean of three housekeeping genes (GAPDH, 16s, EFα1). Error bars show the standard deviation. Significant differences (LSD): P < 0.05 (*); P < 0.01(**).

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Fig 6 Expand

Fig 7.

Time series of gonad developmental stages in Calanus sinicus in different temperatures by food conditions.

(a) 9NF (9°C not fed), (b) 18NF (18°C not fed), (c) 9F (9°C fed) and (d) 18F (18°C fed). GS1 to GS4 represent gonad development stages 1–4.

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Fig 7 Expand

Fig 8.

Gonad development states (GS2-GS4) in Calanus sinicus at the end of the culture experiment.

(a) Group 9NF (9°C not fed), GS2, developing oocytes (OS1 and OS2); (b) Group 9F (9°C fed), GS4, mature oocytes (OS4); (c) Group 18NF (18°C not fed); (d) Group 18F (18°C fed). Scale bar: 400 μm.

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Fig 9.

Relative gene expression variation of gene Vitellogenin receptor (VgR) of Calanus sinicus.

9NF = 9°C not fed, 9F = 9°C fed, 18NF = 18°C not fed and 18F = 18°C fed. D0 to D16 represent Days 0 to 16. The expression level was normalized to the geometric mean of three housekeeping genes (GAPDH, 16s, EFα1). Error bars show the standard deviation. Significant differences (LSD): P < 0.05 (*); P < 0.01(**).

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Fig 10.

Oil sac proportion (OSV%) of Calanus sinicus cultured in different temperatures by food conditions.

9NF = 9°C not fed, 9F = 9°C fed, 18NF = 18°C not fed and 18F = 18°C fed.

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Fig 11.

Cumulative lipid costs for respiration and total lipid loss in oil sacs of Calanus sinicus.

(a) 9NF (9°C not fed), (b) 18NF (18°C not fed), (c) 9F (9°C fed) and (d) 18F (18°C fed).

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Fig 12.

Relative gene expression variation of genes associated with lipid consumption of Calanus sinicus.

(a) HOAD (hydroxyacyl CoA dehydrogenase) expression. (b) DI (dienoyl-CoA isomerase) expression. 9NF = 9°C not fed, 9F = 9°C fed, 18NF = 18°C not fed and 18F = 18°C fed. D0 to D16 represent Days 0 to 16. The expression level was normalized to the geometric mean of three housekeeping genes (GAPDH, 16s, EFα1). Error bars show the standard deviations. Significant differences (LSD); P < 0.05 (*), P < 0.01(**).

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Fig 12 Expand

Fig 13.

Relative gene expression variation of gene ferritin of Calanus sinicus.

9NF = 9°C not fed, 9F = 9°C fed, 18NF = 18°C not fed and 18F = 18°C fed. D0 to D16 represent Days 0 to 16. The expression level was normalized to the geometric mean of three housekeeping genes (GAPDH, 16s, EFα1). Error bars show the standard deviations. Significant differences: LSD, p <0.05 (*), p <0.01(**).

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Fig 14.

Principal component analysis of the relative gene expression data of Calanus sinicus in Group 18F (18°C fed).

(a) The loading plot of six genes. (b) Scatterplot of the averages of the first two principal component scores for C. sinicus. D0 to 16 represent Days 0 to 16.

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Fig 15.

Termination and subsequent development of the over-summering Calanus sinicus cultured in Group 18F (18°C fed).

The internal processes were divided into four periods: quiescence (Day 0), termination and molting process (Days 1–6), early gonad development (Days 6–9) and final gonad development (Days 9–16). The associated metabolic rates (OCR, dotted line), lipid reserve (OSV%, solid line) and gene expression patterns are shown. See Table 1 for gene abbreviations.

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Fig 15 Expand