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Fig 1.

A map of all platforms surveyed by ROV during this study.

Platform code and position shown. Point of first sighting (2007) shown as a triangle (GI-93C).

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Table 1.

A list of all platforms surveyed by ROV during this study.

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Fig 2.

Percent wins in competition for space between Tubastraea micranthus and all other sessile epifauna pooled, by platform.

Percent competitive success plus 95% confidence limits shown. The competition variable is defined as the number of competitive successes over the total number of interactions within a taxonomic group. An overgrowth frequency of significantly >50% was considered to be an indicator of competitive success. A total of 15 platforms were surveyed. Those platforms possessing a number of interactions sufficient for quantitative analysis were included in the study. Data tested for significant variation from 1:1 ratio of competitive successes to losses. Range of number of interactions per platform (n): 18–361. Overall competitive success was significantly higher than the expected 50% level over all platforms (p < 0.01, Fisher’s Exact Test). Highly variable win frequencies between platforms (p < 0.001, Goodness of Fit Test, G-statistic). No significant sub-sets of platforms.

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Fig 3.

Percent success in competition for space between Tubastraea coccinea and all other sessile epifauna pooled, presented by platform.

Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 217–1,283. See Fig 2 legend for additional details. Overall competitive success significantly higher than 50% (p < 0.001, Fisher’s Exact Test). Also, almost all platforms exhibited a competitive win frequency significantly higher than 50% (p < 0.01–0.001, Goodness of Fit Test, G-statistic), except ST-81A and ST-185B, which exhibited no significant difference from the 50% competitive success level (p > 0.05).

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Fig 4.

Percent success in competition for space between Tubastraea micranthus vs. T. coccinea.

Here, the former species is the target organism. Presented by platform. Mean plus 95% confidence limits shown. See Fig 2 legend for additional details. No significant difference in overall competitive success (p > 0.05, Fisher’s Exact Test).

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Fig 5.

Percent of successes in competition for space between Tubastraea coccinea vs. T. micranthus.

Here, T. coccinea was the target organism. Data presented by platform. Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 6–155. See Fig 2 legend for additional details. Overall competitive success significantly greater than 50% (p < 0.001, Fisher’s Exact Test). Significantly higher frequency of competitive successes on GI-93C than on other platforms (p < 0.001, Goodness of Fit Test, G-statistic) where competitive success frequency did not differ significantly from a 50% competitive success level (p>0.05

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Fig 6.

Photographs of the six most abundant competitors of space against Tubastraea coccinea on oil/gas platforms in the study region, northern Gulf of Mexico.

Photos are still-captures from an ROV video. a = Xestospongia sp. with commensal zoanthids Parazoanthus catenularis; b = Dictyonella funicularis; c = Haliclona vansoesti; d = Xestospongia carbonaria; e = Mycale carmigropila; f = Phorbas amaranthus.

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Fig 7.

Percent of success in competition for space between Tubastraea coccinea vs. Xestospongia sp. (with commensal zoanthids Parazoanthus catenularis), by platform.

Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 19–74. See Fig 2 legend for additional details. Overall competitive success significantly higher than the expected 50% (p < 0.001, Fisher’s Exact Test). Also, almost all platforms exhibited significantly high frequencies of competitive success (p < 0.05–0.001, Goodness of Fit Test, G-statistic). Only coral populations on MC-311A and GI-115A exhibited non-significant levels of competitive success (< 50% frequency of success, p > 0.05).

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Fig 8.

Percent of competitive success in competition for space between Tubastraea coccinea and Dictyonella funicularis, by platform.

Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 13–83. See Fig 2 legend for additional details. Overall competitive successes significantly higher than 50% (p < 0.001, Fisher’s Exact Test). Highly variable win frequencies between platforms. Platforms exhibiting significant competitive success were GI-90A, MC-311A, GI-116A, MC-109A, SP-87D, and ST-185A (p < 0.01–0.001, Goodness of Fit Test, G-statistic). The remainder exhibited no significant difference with a 50% competitive success level (p > 0.05).

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Fig 9.

Percent of successes in competition for space between Tubastraea coccinea and Haliclona vansoesti, by platform.

Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 16–60. See Fig 2 legend for additional details. No significant difference between overall observed frequency of competitive success, and an expected 1:1 ratio of the two competitors (p > 0.05, Fisher’s Exact Test). Highly variable success frequencies between platforms. T. coccinea populations on MC-311A, GI-115A, GI-116A, ST-185A, SP-89B, SP-87D, and ST-206A exhibited significant competitive success over this sponge (p < 0.05–0.01, Goodness of Fit Test, G-statistic). The GI-94B coral population exhibited significant competitive losses (p < 0.01). GI-93C, GI-90A, and MC-109A showed no significant variance from the expected 50% competitive success level (p >0.05).

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Fig 10.

Percent of success in competition for space between Tubastraea coccinea and Xestospongia carbonaria, by platform.

Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 20–58. See Fig 2 legend for additional details. Overall competitive success significantly higher than 50% (p < 0.001, Fisher’s Exact Test). Highly variable success frequencies between platforms. T. coccinea on GI-93C, GI-90A, MC-311A, GI-115A, GI-116A, ST-206A, and ST-185B all exhibited significantly high competitive success (p < 0.01–001, Goodness of Fit Test, G-statistic). The remainder exhibited competitive success frequencies which did notvary significantly from 50% (p> 0.05).

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Fig 11.

Percent success in competition for space between Tubastraea coccinea and Mycale carmigropila, by platform.

Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 13–150. See Fig 2 legend for additional details. Overall competitive success significantly higher than 50% (p < 0.001, Fisher’s Exact Test, Goodness of Fit Test, G-statistic). T. coccinea populations were highly variable with respect to competitive success for this interaction. T. coccinea on GI-93C, GI-90A, GI-115A, GI-116A, SP-89B, and ST-185A all exhibited significantly high competitive success (p < 0.01–001, Goodness of Fit Test, G-statistic). Populations on the remainder of the platforms exhibited frequency of success not significantly different from 50% (p> 0.05).

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Fig 12.

Percent of wins in competition for space between Tubastraea coccinea and Phorbas amaranthus, by platform.

Mean plus 95% confidence limits shown. Range of number of interactions per platform (n): 17–150. See Fig 2 legend for additional details. Overall competitive success significantly higher than the 50% level (p < 0.01, Fisher’s Exact Test). Frequency of competitive success varied significantly between platforms. The following platforms exhibited significantly high competitive success frequencies: GI-93C, GI-90A, GI-116A, MC-109A, and SP89B (p < 0.05–0.01, Goodness of Fit Test, G-statistic). The remainder (most of the platforms) exhibited competitive success frequencies not significantly different from 50% (p > 0.05).

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