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Figure 1.

Uredinia and urediniospores of Pst are parasitized by Cladosporium cladosporioides.

A: A uredinium infected by C. cladosporioides (×800), B: A non-infected uredinium filled with urediniospores (×700), C: Hyphae of C. cladosporioides growing from or between urediniospores at an early stage of infection (×2,000), note the shriveled urediniospores, D: Numerous hyphae of C. cladosporioides growing on a uredinium at a late stage of infection (×400), note presence of very few normal urediniospores, E: Urediniospores invaded by C. cladosporioides hyphae (×1,800), F: High magnification of a C. cladosporioides hypha invading a urediniospore (×11,000).

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Figure 1 Expand

Figure 2.

Morphological features of Cladosporium cladosporioides cultured on MEA medium.

A: Fungal colonies on MEA medium, B: Branched ramoconidia and chains of conidia, C: Ramoconidia and dense conidia on the tip of a conidiophore, D: Oval to spherical conidia and separated ramoconidia with a dark scar at one end or both ends.

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Figure 3.

Morphological features of Cladosporium cladosporioides under a scanning electron microscope.

A: Conidial chains (×7,000), B: Scars on a conidiophore (×20,000), C: Top view of a conidiophore with scars (×7,000), D: scars on a secondary ramoconidium (×4,000), E: Conidia on conidiophores (×5,000), F: Hyphae (×2,000).

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Figure 4.

The pathogenicity test to confirm that C. cladosporioides could hyperparasite Pst.

A, B: CK1, wheat leaves (Cultivar: MingXian169) inoculated with C. cladosporioides, 14 dpi, without symptoms; C, D: CK2, Symptoms on wheat leaves (MingXian169) inoculated with CYR32, 20 dpi; E, F, G: Pustules on wheat leaves (same cultivar and race as above) inoculated at 14 dpi with C. cladosporioides suspension. Symptoms at 5d, 7d and 9d later.

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Figure 5.

SEM observations of Pst inoculated with C. cladosporioides.

A: 12 h after inoculation, the spore of C. cladosporioides contacted the urediniospore surface (×7,000), B: 24 h after inoculation, spores of C. cladosporioides germinated (×700), C: 36 h after inoculation, direct penetration by germ tube of C. cladosporioides into a urediniospore (×10,000), D: 48 h after inoculation, amount of urediniospores were surrounded with germinated spores of C. cladosporioides (×2,000), E: 72 h after inoculation, germ tubes of C. cladosporioides have penetrated urediniospores (×3,000), F: 120 h after inoculation, C. cladosporioides has colonized the urediniospores (×1,300).

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Figure 6.

A neighbor-joining tree constructed using the sequences of ITS,EF and ACT to determine the phylogenetic relationship of the hyperparasitic fungal strain CGMCC 7.175 (bold and marked with *) isolated from uredinia of Pst with other accessions of Cladosporium spp. in GenBank.

The values at braches are percentages from 1,000 bootstrap replications.

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Table 1.

Accession numbers of Cladosporium species used in the molecular characterization.

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Table 1 Expand