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Figure 1.

Distribution of gray wolf dens detected between 2011 and 2012 in Hamedan province, Iran.

Wolf dens were overviewed in a context of topography and main roads in Hamedan province, Iran.

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Figure 1 Expand

Figure 2.

Proportion of land use/land cover categories used in this study.

Proportion of each cover type was calculated within 2 km circular buffers around wolf den sites (den areas), random locations (random areas) and proportion of the whole study area (Hamedan province).

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Figure 2 Expand

Table 1.

Mean (SE) values of variables measured at the level of the den area, in 2 km circular buffers with and without wolf dens in Hamedan province, Iran.

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Table 1 Expand

Table 2.

Mean (SE) values of fine-scale variables measured in sample plots with and without wolf dens in Hamedan province, Iran.

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Table 2 Expand

Table 3.

Selected candidate Generalized Linear Models explaining gray wolf den area selection patterns in Hamedan province, Iran, at the landscape level.

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Table 3 Expand

Table 4.

Relative importance (W+), model-averaged coefficient estimates (Estimate), and unconditional standard errors (SE) for the predictors included in the selected candidate models determining the probability of a given area being selected as a den area by wolves in Hamedan province, Iran (models with ΔAICc <2).

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Table 4 Expand

Table 5.

Selected candidate Generalized Linear Models explaining gray wolf den site selection patterns in Hamedan province, Iran, at the fine spatial scale.

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Table 5 Expand

Table 6.

Relative importance (W+), model-averaged coefficient estimates (Estimate), and unconditional standard errors (SE) for the predictors included in the selected candidate models determining the probability of a given site being selected as a den site by wolves in Hamedan province, Iran (models with ΔAICc<2).

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Table 6 Expand