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Figure 1.

Macroscopic appearance of biopsied lesions from acne patients.

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Table 1.

Clinical characteristics of acne patient group from Finland.

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Table 2.

Clinical characteristics of acne patient group from Germany.

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Figure 2.

2-components NMF (Non Negative Matrix Factorization) model.

The heatmap of 904 regulated genes on subjects-corrected log2-expressions (legend color blue for low expression and red for high expression).

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Figure 3.

Bioinformatic analysis of differentially modulated genes.

Gene enrichment analysis in three different functional ontologies were performed using MetaCore: biological processes (a), process networks (b) and canonical pathway maps (c).

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Table 3.

Top 20 up-regulated genes in the German acne patient group based on Affymetrix microarray technology.

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Table 4.

Top 20 down-regulated genes in the German acne patient group based on Affymetrix microarray technology.

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Figure 4.

Th17 related cytokines were increased in acne lesions.

The skin biopsies were obtained from healthy volunteers normal skin (NS), from acne patients non-lesional skin (A NS) and lesional skin (AL) and from psoriasis patients non-lesional skin (P NS) and lesional skin (PL). The expression levels of cytokines were determined by RT-PCR. The expressions of inflammatory cytokines TGF-β, IL-6, IL-1β and IL-23 were increased significantly both in acne and psoriasis but IL-1β and TGF-β expression was higher in acne. IL-23 was increased more in psoriasis lesions. IL-17A was significantly expressed in lesional acne and psoriasis skin. IL-22, was slightly elevated in acne and psoriasis, however significantly more in psoriasis. *P<0.05, **P<0.01, ***P<0.001; bars represent mean +SEM. RU, relative units; TGF-β, transforming growth factor- beta; Th17, T helper type 17.

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Table 5.

Expression at the mRNA level of markers and cytokines characterizing the Th17 cell subtype in the German acne cohort.

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Figure 5.

Inflammatory cell infiltrates of early acne lesions in immunohistochemistry.

Representative immunohistochemical staining showing IL-17A+ cells in early acne lesion (a). Staining of IL-17A+ and CD3+ cells shows that IL-17A (in green) and CD3 (in red) were detected in acne vulgaris lesions. Cell nuclei were counterstained in blue. Arrows indicate IL-17A and CD3 double positive cells (b). T-bet+ cells (c) were more numerous than IL-17A+ cells or Foxp3+ cells (d) in the lesional acne skin. Large number of CD4+ cells (e), mostly lymphocytes, was seen around pilosebaceous unit and perivascularly. CD8+ cells (f) were fewer in number than CD4+ cells. Large number of CD68+ macrophages (g) and a few CD83+ cells (h), which are mature dendritic cells, were detected around sebaceous follicles. (Bar = 200 µm; in the insets bar = 50 µm).

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Table 6.

Expression at the protein level of markers and cytokines characterizing the Th17 cell subtype in the German acne cohort.

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Table 7.

Number of positively stained cells in lesional and non-lesional skin of acne patients (n = 16)1 as assessed by immunohistochemistry.

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Figure 6.

The expression of proinflammatory mediators in Finnish patient cohort examined by RT-PCR.

The expression of a) toll-like receptors TLR2 and TLR4, b) proinflammatory cytokine TNF-α and neutrophil chemotactic factor IL-8 were significantly up-regulated in lesional acne. c) IL-20 mRNA were increased in acne and psoriasis lesions. It is known that IL-20 can stimulate epidermal hyperplasia in model systems and is upregulated in psoriasis. *P<0.05, **P<0.01, ***P<0.001; bars represent mean ±SEM.

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Table 8.

Expression at the mRNA level of proinflammatory mediators in the German acne cohort determined by Affymetrix microarray technology.

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Figure 7.

Antimicrobial peptides are increased in acne and psoriasis lesions by RT-PCR.

The expressions of mRNAs were significantly higher in psoriasis except for hCAP-18, which was similarly expressed in both. *P<0.05, **P<0.01, ***P<0.001; bars represent mean ±SEM. BD2, human beta-defensin 2; BD3, human beta-defensin 3; hCAP-18, human cathelicidin antimicrobial protein 18; LCN2, lipocalin-2; RU, relative units.

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Table 9.

Expression at the mRNA level of antimicrobial peptides in the German acne cohort determined by Affymetrix microarray technology.

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Figure 8.

The expression of Th1 cell and Treg markers were activated in acne by RT-PCR.

Th1 transcription factor T-bet, Th1 cell surface chemokine receptor CXCR3 and IFN-γ (the effector cytokine of Th1 cells) mRNAs were significantly up-regulated in acne and psoriasis lesions. The transcription factor of Tregs, Foxp3, was elevated in acne and psoriasis. TGF-β which is both Th17 and Treg lineages inducing cytokine was significantly increased in acne lesions. The product of Tregs, immunosuppressive cytokine IL-10 was expressed more in acne than in psoriasis. *P<0.05, **P<0.01, ***P<0.001; bars represent mean ±SEM. RU, relative units; TGF-β, transforming growth factor- beta; Th1, T helper type 1; Treg, regulatory T cell.

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Table 10.

Expression at the mRNA level of markers and cytokines characterizing the Th1 cell subtype in the German acne cohort determined by Affymetrix microarray technology.

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Table 11.

Expression at the mRNA level of markers and cytokines characterizing the Treg cell subtype in the German acne cohort determined by Affymetrix microarray technology.

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Figure 9.

The effect of IL-17 on keratinocytes.

Th17-cells are defined to secrete IL-17 cytokines. Also some other cells can secrete IL-17 cytokines when properly activated. IL-17A and IL-17F regulate genes on keratinocytes, that are involved in innate immune defence, such as antimicrobial peptides (beta-defensins, S100 proteins A7/9, cathelicidin, LCN2) and a range of chemokines G-CSF, CXCLs and CCL20 which regulate neutrophil and lymphocyte trafficking. IL-17A and IL-17F stimulate keratinocytes leading to keratinocyte proliferation. Also vascular inflammation is activated.

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