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Figure 1.

An example of the delineation of habitat patches at one BBS location.

A two-dimensional vegetation map (A) and a vegetation map segmented by height structure (B) are shown. The pixel-based segmentation method (supporting information S1) is used to segment two dimensional habitat maps by using height thresholds.

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Figure 2.

Distribution of BBS routes through the primarily forested ecoregions in the U.S.

The richness models for the woodland guild were built using data from both eastern and western forested ecoregions. The forest edge and interior forest bird richness was modeled in the eastern forested ecoregions only.

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Figure 2 Expand

Table 1.

List of all metrics developed in the study.

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Table 1 Expand

Figure 3.

Guild richness associations with various metrics.

(Top row): correlation bar plots of the most predictive metrics of species richness by guild. White bars represent a positive correlation and grey indicate a negative correlation. (Bottom rows): correlation comparisons between comparable patch-based metrics with and without considering the vertical patches and edges for the woodland and forest edge guild. The left panels show traditional metrics without accounting for height-heterogeneity; the right panels are height-structured counterparts. The black dots indicate a negative correlation and the grey ones indicate a positive correlation.

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Figure 4.

Predictive ability of multivariable models.

A, B, C, and D are the four habitat metric sets, and 4BPHMs are the four best predictive height-structured metrics. Each of the top panels shows four linear models with whiskers giving 95% confidence interval of adjusted-r2 values. The length of the bar represents the mean adjusted-r2 for these models. The lower panels show the explained variance of the comparable random forest (RF) models. Uniquely the top bars at lower pannels are the results from the models employed all metrics from the four metric sets.

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Figure 5.

Random Forest model results.

(Top row): Modeled vs. actual species richness for three guilds using all-inclusive random forest models. (Below the scatter plots): variable importance plots show the percent increase in mean square error (%IncMSE) of the top 20 most influential metrics in the woodland guild richness model and the forest edge guild richness model (note different scales on X-axes). The metrics characterizing vegetation height heterogeneity are plotted with triangles and the rest of the metrics are circles.

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