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Figure 1.

Tilivalline structure, chemical formula, and mass.

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Table 1.

Sources and origin of Klebsiella oxytoca used in this study.

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Table 2.

Antimicrobial resistance of K. oxytoca.

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Figure 2.

Animal isolates of K. oxytoca produce cytotoxin.

HEp-2 cell culture inoculated with K. oxytoca supernatant and media control at 4, 10, and 30× magnifications. A) Media control showing normal morphology of cells stained with Diff-quick after 48 hours of incubation. B) K. oxytoca, ATCC 13182, isolate negative for cytotoxin production showing normal cell morphology. C) K. oxytoca isolate, 09-7231-1, positive for cytotoxin production showing abnormal cell morphology, decreased concentration of attached cells, small round cells, and multinucleated cells.

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Table 3.

Cytotoxin of K. oxytoca varies with different growth conditions.

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Figure 3.

Temporal pattern of cytotoxicity in various conventional bacterial growth media.

K. oxytoca, 09-7231-1, was cultured in A) LB; B) TSB; and C) HBI liquid media. At various time points, the supernatant was collected and used in the cytotoxicity assay using HEp-2 cells. TSB media has the strongest cytotoxin induction capability compared to LB and HBI media. The degree of cytoxicity is indirectly proportional to the level of monolayer confluency i.e., the higher % the confluency, the lower the cytotoxity and vice versa.

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Figure 4.

Soy components in TSB induced HEp-2 cell detachment and death.

HEp-2 cells treated with A) media containing only casein extract (Tryptone; control); B) supernatant of K. oxytoca, 09-7231-1, grown in media containing only casein extract; C) media containing only soy extract (Soytone); D) supernatant of K. oxytoca, 09-7231-1, grown in the media containing only soy extract.

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Figure 5.

Tilivalline induced HEp-2 cell detachment and death.

A) HEp-2 cells incubated with the control DMSO, which was used to dissolve tilivalline. The majority of cells adhered to the bottom of the cell culture well after 48 hours of culture; B) HEp-2 cells treated with purified tilivalline (1 µg/ml) were detached from the cell culture well after 48 hours of incubation.

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Figure 6.

The reference proteins encoded by CTA, CFX B, and PAX B are displayed as colored bars.

The arrow beneath the bar shows the region in the reference protein to which the corresponding K. Oxytoca protein aligns from the BLAST analysis.

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