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Figure 1.

Distribution of habitat types across the major clades of water scavenger beetles.

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Figure 2.

A selection of habitat diversity within the Hydrophilidae.

A) Lentic habitat in a vegetated marsh, B) Lotic habitat along a mountain stream, C) Coastal dune pond, D) Detrital forest pool, E) Anacaena sp. (Chaetarthriinae) inside a drained bromeliad tank, F) Oocyclus sp. (Laccobiini: Laccobius group) on a wall seepage, G) Pelosoma sp. (Megasternini) emerging from a Heliconia inflorescence, H) wet sand habitat along a creek, I) Dactylosternum sp. (Coelostomatini) under the bark of a freshly cut tree, J) Chimaearocyon shimadai (Megasternini) in the brood chamber of Pheidole ants, photo credit: Taku Shimada, K) Rygmodus sp. (Rygmodinae) on flowers, photo credit Richard Leschen, L) Nitidulodes sp. (Megasternini) on an aroid inflorescence. All photos by A.E.Z. Short unless otherwise indicated.

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Figure 3.

Time-calibrated phylogeny for major lineages of Hydrophilidae, along with estimated species richness values.

The star indicates the location of the diversification rate increase determined by our MEDUSA analysis.

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Figure 4.

Relationship between clade age and estimated species richness in Hydrophilidae.

The shaded regions show 95% confidence interval of the expected diversity under a low (e = 0, solid lines) and high (e = 0.9, dashed lines) extinction.

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Figure 5.

Ancestral character reconstructions of habitat transitions on the time-calibrated phylogeny of Hydrophilidae.

The pie charts at nodes indicate maximum likelihood support for ancestral states. A yellow bar indicates the location of the diversification rate shift determined by MEDUSA.

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