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Figure 1.

Transect outline.

Transects were replicated 20 times along each of the 3 roads. Each plot (road, mid, far) measured 2×50 m, with the roadside plot parallel to the road, and the mid and far plots perpendicular. Mid and far plots both are divided in two subplots of 2×25 m each (mid1, mid2, far1 and far2).

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Figure 2.

Native species richness as a function of elevation and distance to the road.

(A) Average native species richness (±1 SE) in plots across the elevational gradient. Different letters indicate significant differences (p<0.05) in a Tukey’s post-hoc test. (B) Native species richness (number of species per plot) as a function of elevation. •, black full line: roadside plots; •, grey full line: mid plots; ○, broken line: far plots (see Fig. 1 for plot types).

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Figure 3.

Plot coverage as a function of distance to the road.

Percentage of the plots (± 1 SE) covered with (A) total vegetation, (B) bryophytes, (C) the dominant species (E. nigrum) and (D) bare ground, as a function of distance to the road. Different letters indicate significant differences (p<0.001).

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Figure 4.

Ordination of plots.

DCA-ordination of plots (•) based on total species coverage. Ellipses indicate the standard deviations for different subgroups as a function of elevation and distance to the road. Elevation: H = highest third, M = middle third, L = lowest third of the gradient; road distance: A = roadside, B = mid, C = far (see Fig. 1 for plot types). The arrow represents the vector of increasing elevation. Eigenvalues of DCA1 and 2 are 0.3479 and 0.2771 respectively.

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Figure 5.

Changes in total species composition between roadside and far plots with elevation.

Left column: number of species that were present in the natural plant communities but lost in the roadsides ( = species unique for the natural plant communities). Middle column: species newly gained in the roadsides ( = species unique for the roadsides). Right column: shared species between roadsides and natural plant communities. A-C = all species, D-F = competitive species, G-I = stress-tolerant species, J-L = ruderals, derived from Grime’s triangle. Significance of linear regressions: ***: p<0.001; **: p<0.01; *: p<0.05;.: p<0.1; otherwise higher than 0.1.

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Figure 6.

Alien species richness as a function of elevation and distance to the road.

(A) Alien species richness (number of species per plot) as a function of elevation. Roadside plots (•, black line), intermediate plots (•, grey line) and far plots (○, dashed grey line). (B) Ratio of alien species richness in the natural plant communities to that in the roadside plot, with mid/roadside (•, grey line) and far/roadside (○, dashed grey line). Significance of linear regressions: see text. Symbols of different variables were slightly shifted to avoid overlap.

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Figure 7.

Elevational range of native and alien species.

Relationship between elevational range and maximum elevation reached by native (•, dashed line) and alien (○, thick solid line) species. Symbols are constrained to the lower right of the graph (marked by the thin solid line), since the elevational range cannot exceed the maximum elevation.

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