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Figure 1.

Cell model.

The cell membrane acts as a capacitance, which is submitted to the membrane potential . Three blocks exchange ions between the cytosol and the outer medium: they induce electroneutral transports (with the and exchangers), electrogenic currents via the Na/K-ATPase, and passive ion channels (for , , and ). The chemical reactions are assumed to take place within an homogenous cytosol.

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Figure 1 Expand

Figure 2.

Three dimensional representation of steady-state pH.

pH is drawn as a function of the two controlling parameters and . The flattest area of this surface stretches over units around physiological pH (7.2, in black). This corresponds to values that can be reached by cellular systems (red boundaries).

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Figure 2 Expand

Figure 3.

pH modification by changes in biochemical constants.

The reference values are , , , and . For and modification, the ratio is held constant (see text for the explanation).

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Figure 3 Expand

Figure 4.

Forced acidosis by a simulated spike.

The rise and the decrease of are highlighted by blue areas. (A) The expected pH overshoot takes place with or without the presence of a 60 mM physiological buffer (dashed lines). (B) The ionic ratios relative to the initial values are reported as well.

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Figure 4 Expand

Figure 5.

Simulated ischemia.

Lactate production occurs with a constant , a buffer, and with transporters working at their nominal level. The lactate accumulation is highlighted by a green area. (A) The predicted pH overshoot takes place during the lactate removal by the monocarboxylate transporters. (B) The corresponding ionic ratios relative to the initial values.

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Figure 5 Expand