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Figure 1.

Identification of mNSCs in vitro.

(A) Dumbbell-shaped mitotic cells were identified 72 hours after plating mesencephalic cells from E14 rats. (B) By 4 days in vitro (DIV), neurospheres had grown in size and detached from the culture substrate. (C) A floating 7 DIV neurosphere. (D) Fluorescence photomicrogragh showing a neurosphere at 7 DIV immunoreactive for the stem cell marker nestin. Scale bar = 100 µm. Green: FITC.

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Figure 2.

Identification of fetal DA neurons in vitro.

Fetal DA neurons were isolated from E18 rat ventral midbrain. (A) Phase-contrast photomicrograph showing cells beginning to elaborate neural processes at 4 DIV. (B) With time in culture, neurites grew longer and thicker. (C, D) Immunofluorescence staining at 9 DIV showing TH-immunopositive neurons with a variety of distinct morphologies (unipolar, bipolar, and multipolar). Green: FITC. (E, F) Fluorescence photomicrograph showing that many cells express both the neuronal marker β-III-tubulin and the DA marker TH. Green: FITC; Red: TRITC. (G) DAPI (blue) nuclear counterstaining; (H) Merged image of E, F, and G. Scale bar = 100 µm.

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Table 1.

Mean rotation index.

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Figure 3.

Identification of mesencephalic neural stem cells overexpressing GDNF (GDNF-mNCSs).

(A) Fluorescence photomicrographs show that all neurospheres were immunoreactive for the NSCs marker nestin. The neurospheres of peGFPN1-GDNF-transfected mNSCs (termed GDNF-mNSCs) were strong immunopositive for GDNF compare to mNSCs transfected with peGFPN1. Green: FITC, Red: TRITC. Scale bar = 50 µm. (B) Prussian blue staining and transmission electron microscopy confirmed SPIO particles in the cytoplasm of mNSCs. (C) GDNF mRNA expression in GDNF-mNSCs compared to mNSCs transfected with peGFPN1 (termed control mNSCs) was determined by RT-PCR. β-actin was used as an internal control. (D) GDNF protein expression in GDNF-mNSCs and control mNSCs was determined by Western blot. Relative protein expression was normalized to β-actin. The data represents the mean ± SD. *P<0.05 vs. control mNSCs. peGFPN1-GDNF-transfected mNSCs (termed GDNF-mNSCs); mNSCs expressing empty vector (termed control mNSCs).

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Figure 4.

In vitro differentiation of SPIO, eGFP double-labeled GDNF-mNSCs.

Fluorescence photomicrographs showing SPIO, eGFP double-labeled GDNF-NSCs growing in a differentiation medium for seven days. eGFP-positive cells (A, D, G, J), immunoreactive for the neuronal markers β-III-tubulin (B) and TH (E), the astrocyte marker GFAP (H), and GDNF (K). Images are merged in the last column (C, F, I, L). Scale bar = 100 µm. Green: FITC, Red: TRITC.

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Figure 5.

MR images (T2W/FFE) showing the survival of SPIO-labeled mNSCs in the striatum.

Note the high density of SPIO-labeled mNSCs in each group at the site of injection (striatum), but no detectable extrastriatal signal.

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Figure 6.

Survival and differentiation of transplanted cells in the striatum.

Representative photomicrograph illustrating that many transplanted cells survived in the striatum. There were more eGFP+ (green), nestin+ (red), nestin+/eGFP+(merged yellow), TH+ (red), and TH+/eGFP+ cells (merged yellow) in the GDNF-mNSCs+DA group than in the other groups. Scale bar = 100 µm. Green: FITC, Red: TRITC.

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Table 2.

Number of cells with specific expression phenotypes and DA neuron differentiation rate after cell transplantation into the rat striatum.

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