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Figure 1.

β-galactosidase staining (Blue) of urinary bladders after intravesical treatment of Adeno-Cre or Adeno-empty vector.

Adeno-Cre (A-C, F) or Adeno-empty vector virus (E) (8.46×108 pfu per mouse) was delivered into the bladder lumen of ROSA mice (A-C, E) or C57 mice (F) for 2 hours. The mice were sacrificed at days 3 (A), 7 (B), or 14 (C) post Adeno-Cre bladder instillation. The bladders were removed and stained for β-galactosidase (blue) expression (counterstaining with fast red). A dynamic change of β-galactosidase is seen with the maximum expression at 7 days post Adeno-Cre instillation (B), predominantly in the upper (inner) 2/3 layer of the urothelium. β-galactosidase activity was reduced when half of the Adeno-Cre dose (4.23 ×108 pfu) (D) was used as compared with the full dose (B). Instillation of either Adeno-Cre-empty into ROSA mice (E) or Adeno-Cre to C57 mice (F) did not show β-galactosidase expression. n=5 for each group. All figures are at 10x magnification.

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Figure 2.

β-galactosidase staining of heart, liver, kidney and lung after intravesical treatment of Adeno-Cre.

Adeno-Cre (8.46×108 pfu per mouse) was delivered into the bladder lumen of ROSA mice for 2 hours (n=5). At day 7, mice were sacrificed, followed by removal of heart (A), liver (B), kidney (C) and lung (D). X-Gal staining for β-galactosidase activity was performed in all these tissues. No observable staining (blue color) was observed (only counterstaining with fast red was present), as compared with Figure 1 (A-D). All figures are at 10x magnification.

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Figure 3.

Immunofluorescent staining of β-galactosideasae and cytokeratin expression of urinary bladder after intravesical treatment with Adeno-Cre.

Adeno-Cre (8.46×108 pfu per mouse) was delivered into the bladder lumen of ROSA mice for 2 hours (n=5). Mice were sacrificed and their bladders were removed and processed for immunofluorescence staining. The urothelial mucosa shows in green β-galactosidase (A) and in red cytokeratin 7 expressions (B). In Figure (C), overlapping of the fields (A) and (B) demonstrates the co-localization of β-galactosidase and cytokeratin 7 expressions (orange-yellow). Nuclei were stained with DAPI (blue color). β-galactosidase expression is scattered and observed mainly in the 2/3 upper (inner) layer of the urothelium versus cytokeratin expression which is evenly present in the full thickness of the urothelium.

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Figure 4.

Breeding scheme and genotyping of double transgenic mice.

Two-step plan for breeding double transgenic mice. Upper: Scheme for breeding plan starting from KrasLSLG12D/+. p53+/+ and Kras+/+. p53fl/fl. Useful mice are marked in blue text. Lower: Agarose gel imaging for genotyping: A double transgenic mouse (mouse 4), Kras LSLG12D/+ /p53 Fl/Fl was characterized by genotyping. Kras genotyping (left): Wild type (WT) = ~ 507 bp; Mutant = ~600 bp (Heterozygote = ~507 bp and ~ 600 bp); p53 genotyping (right): Wild type = ~ 270 bp; Homozygote = ~ 390 bp (Heterozygote = ~ 270 bp and ~ 390 bp).

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Figure 5.

MRI (A) and histology (B-F) images from mice with Adeno-Cre treatment in the presence of sutures.

Kras LSLG12D/+ /p53 Fl/Fl Kras LSLG12D/+ /p53 Fl/Fl mice were intravesically treated with 8.46×108 pfu Adeno-Cre with a distal urethral suture placed for 2 hours, as described in Materials and Methods. When the pelvic tumors appeared (four to eight weeks after Adeno-Cre treatment), proton density-weighted MRI scans were performed and then the mice were sacrificed for histological examinations. A: MR images (coronal plane, left; sagittal plane, right) of lower body of mouse with lower pelvic tumor; B: Low power (2x) H&E section shows that the tumor is located in the pelvic soft tissue but not within the urinary track; sarcoma tumor with focal necrosis is apparent in C (10x); increased mitotic rate is apparent in D (40x); invasion of adipose tissue is apparent in E (40x); and invasion of striated muscle F (40x).

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Figure 6.

Immunohistochemistry of the pelvic soft-tissue sarcoma tumor induced in the Adeno-Cre treated Kras LSLG12D/+ /p53 FL/FL mice.

A: The high Ki67 expression in the nuclei of >70% of the tumor cells demonstrates a high proliferative rate (40x); Sarcoma tumor B (10x); C (40x) shows negative staining for pan-cytokeratin, demonstrating a non-epithelial phenotype; in B, only the skin layer is positive, acting as an internal positive stain control.

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Figure 7.

Induction of urothelial hyperplasia.

Kras LSLG12D/+ /p53 Fl/Fl mice were intravesically treated with Adeno-Cre (8.46×108 pfu Adeno-Cre with 3-hour holding of catheter once per week for a total of four weeks and with no suture applied) 4.5 to 6 months after last instillation. Proton density-weighted MR images (axial plane) of the pelvis of a normal (control) mouse (A-left) and a 6-month treated mouse (A-right) showing focal thickening of the urinary bladder wall (arrow).

As compared with histological images from a normal, control mouse (left B, C and D); the H&E histological sections show the presence of mild urothelial hyperplasia at 4.5 months (B-center) and moderate hyperplasia at 6 months (B-right). Ki67 staining shows no increase in nuclear expression in the bladder of a mouse after 4.5 months (C-center) but increased full thickness with nuclear Ki67 expression in the bladder urothelium after 6 months (C-right). COX-2 staining shows a modest increase in the cytoplasmic and peri-nuclear expression in the bladder urothelium at 4.5 months (D-center) and a moderate increase at 6 months (D-right).

Recombination analysis of Kras and p53 genes was performed with both Kras (E-left) and p53 (E-right): Lane 1: wild type mouse; lane 2: Kras LSLG12D/+ /p53 Fl/Fl mouse before Adeno-Cre treatment; lanes 3 and 4: sarcoma tissue from Kras LSLG12D/+ /p53 Fl/Fl mice after Adeno-Cre treatment with suture (Sar1 and Sar2); lanes 5 and 6: bladder tissues from Kras LSLG12D/+ /p53 Fl/Fl mice after Adeno-Cre treatment without suture (Hyp1 and Hyp2).

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