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Table 1.

Allometric fits for behavioral and physiological phenomena used in the calculation of diffusivity.

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Figure 1.

Allometric relations between bodymass M and population density, metabolic rate, mean longevity, daily displacement, home range, and range length.

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Figure 2.

Allometric relations between bodymass M and animal-mediated nutrient diffusivity, Equations [3] and [4] in the main text.

Solid lines are estimates calculated using scaling arguments, dashed lines as a fit to primary data. Circles show diffusivity by way of excreta, crosses show diffusivity by way of bodymass.

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Figure 3.

Diffusion into granitic region of KNP.

Upper panel shows geometry of the simulated transect, with an inset to show the initial and boundary conditions of a transect across the substrate gradient in the absence of herbivore diffusion. Lower panels show phosphorus stocks in edible vegetation under a succession of herbivore removals, varying from Φ varies from 7 km2/year (estimate prior maximum) to 2 (present-day estimate) to 0.075 (estimate in the absence of herbivores >100 kg). A. P dynamics under an upper estimate of K = 0.0013/year; B. P dynamics under a lower estimate of K = 0.00013/year. Additional parameter values set to Po = 875 kg P km−2, G = 0.5 kg P km−2 year−1.

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Figure 4.

Estimates of herbivory and nutrient diffusivity in Kruger National Park by mammalian herbivores >1 kg in KNP.

(a) Potential rates of consumption based on population density and metabolic demand. The mean rate of herbivory per taxon is 927 kg/km2 or 0.37% of the biomass standing crop. (b) cumulative rate of herbivory across body mass (c) nutrient diffusivity Φ, using observations of component terms (where possible – black points) and allometric scaling (8.672*M1.191; red points). (d) cumulative nutrient diffusivity Φ across body mass, based on direct observations of component terms, and allometric scaling of component terms.

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Figure 5.

Global distribution of terms in herbivore diffusion of nutrients.

(a) nutrient diffusivity Φ = DQ/αB, (b) change in Φ if all threatened species are lost.

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Figure 6.

Global distribution of terms in herbivore diffusion of nutrients.

(a) movement diffusivity D, (b) percent consumed biomass Q/αB, (c) total animal biomass (ie Σ mass * population), (d) nutrient diffusivity Φ = DQ/αB, (e) edible biomass αB, (f) number of mammalian herbivores >1 kg.

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