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Figure 1.

Attenuating liver IRI through low-dosage LPS preconditioning.

Mice were subjected to 90 mim of partial liver ischemia, followed by 6 h and 24 h reperfusion. (A) and (B) Hepatocellular function evaluated by ALT (U/L) and LDH (U/L). Mean±SD, **P<0.001 versus sham group; ##P<0.001 versus IR group. (C) Histopathalogic analysis of livers harvested 6 hours after reperfusion: (a) Sham group: Normal hepatic architecture; (b) IR group: severe hepatic lobule distortion, sinusoidal congestion, apparent edema, vacuolization and massive necrosis; (c) LPS PC+IR group: mild vacuolization, punctate necrosis and edeman. (D) The severity of liver IRI by Suzuki’s histological grading.

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Figure 2.

Reduction of hepatocellular apoptosis after liver IR through LPS preconditioning.

(A) Liver apoptosis by TUNEL stainning: (a) sham group; (b) IR group and (c) LPS PC+IR group. (B) Apoptotic cells were quantified in six high-power fields (400×), and expressed as percentages of apoptotic cells among total cells. Mean±SD, **P<0.001 versus sham group; #P<0.05 versus IR group. (C) Caspase-3 activity, Mean±SD, **P<0.001 versus sham group; ##P<0.001 versus IR group.

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Figure 3.

ATF4-CHOP pathway and related apoptotic pathway inhibition by LPS preconditioning after IR.

(A) Western-assisted analysis of ATF4, CHOP, Cleaved Caspase-12, Cleaved Caspase-3 and β-Actin. Representative of three experiments. (B) Relative quantities of protein of ATF4, CHOP, Cleaved Caspase-12, Cleaved Caspase-3 to β-Actin, Mean±SD, **P<0.001 versus sham group; ##P<0.001 versus IR group; #P<0.05 versus IR group. (C) immunohistochmistry analysis of CHOP: (a) sham group; (b) IR group and (c) LPS PC+IR group. Positive cells were quantified in six high-power fields (400×), and expressed as percentages of positive cells among total cells. Mean±SD,**P<0.001 versus sham group; ##P<0.001 versus IR group.

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Figure 3 Expand

Figure 4.

ATF4-CHOP pathway inhibited by low-dose LPS preconditioning in hepatocytes.

(A) Western-assisted analysis of ATF4, CHOP and β-Actin. Representative of three experiments. (B) Relative quantities of protein of ATF4 and CHOP to β-Actin, Mean±SD, **P<0.001 versus sham group; #P<0.05 versus IR group. (C)The released LDH level of hepatocytes after TM or H2O2 treatment, Mean±SD, **P<0.001 versus sham group; #P<0.05 versus IR group.

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Figure 5.

ATF4 knockdown protection against hepatocellular death induced by H2O2 in vitro.

(A) Western-assisted analysis of ATF4, CHOP, Cleaved Caspase-12, Cleaved Caspase-3 and β-Actin. Representative of three experiments. (B) Relative quantities of protein of ATF4, CHOP, Cleaved Caspase-12 and Cleaved Caspase-3 to β-Actin, Mean±SD, **P<0.001 or *P<0.05 versus CONsiRNA. (C)The released LDH level of hepatocytes after H2O2 treatment, **P<0.001 versus CONsiRNA.

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Figure 5 Expand

Figure 6.

Role of ATF4-CHOP pathway in modulating immune responses of LPS preconditioning.

(A) Cytokine gene (TNF-α, IL-6 and IL-10) expression in livers harvested 6 hours after reperfusion by Quantitative RT-PCR analysis. Mean ± SD, **P<0.001 versus sham group; ##P<0.001 versus IR group; #P<0.05 versus IR group. (B) Cytokine gene (TNF-α, IL-6 and IL-10)expression in BM-macrophage stimulated by 1 µg/ml LPS treatment for 3 hours. Mean ± SD, **P<0.001 versus CON group; *P<0.05 versus CON group, ##P<0.001 versus LPS PC+LPS group. (C) Cytokine secretion (TNF-α, IL-6 and IL-10) by ELISA in BM-macrophage stimulated by 1 µg/ml LPS treatment for 24 hours. Mean ± SD, **P<0.001 versus CON group; *P<0.05 versus CON group, ##P<0.001 versus LPS PC+LPS group. (D) Western-assisted analysis of ATF4, CHOP, P-NF-κB p65, IκBα, SCOS3, p-ERK1/2 and β-Actin. Representative of three experiments. The relative quantities of protein to β-Actin, Mean±SD, **P<0.001 versus CON group, #P<0.05 versus LPS PC+LPS group.

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Figure 7.

ATF4 knockdown inhibition of immune response in vitro.

(A) Western-assisted analysis of ATF4, CHOP, P-NF-κB p65, IκBα, SCOS3, p-ERK1/2 and β-Actin. Representative of three experiments. The relative quantities of protein to β-Actin, Mean±SD, **P<0.001 versus CON group. (B) Cytokine secretion (TNF-α, IL-6 and IL-10) by ELISA in BM-macrophage stimulated by 1 µg/ml LPS treatment for 24 hours. Mean ± SD, **P<0.001 or *P<0.05 versus CONsiRNA group.

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Figure 8.

Knockdown of ATF4 improves hepatocellular function and ameliorates liver IRI.

Mice were subjected to 90 mim of partial liver ischemia, followed by 6 h reperfusion. (A) Hepatocellular function evaluated by ALT (U/L). Mean±SD, **P<0.001; ##P<0.001. (B) Histopathalogic analysis: (a) Sham group: normal hepatic architecture; (b) CON group: severe hepatic lobule distortion, sinusoidal congestion, apparent edema, vacuolization and massive necrosis; (c) ATF4 siRNA group: mild vacuolization, punctate necrosis and edeman; and (d) Ns siRNA: severe hepatic lobule distortion, sinusoidal congestion, apparent edema, vacuolization and massive necrosis. (C) The severity of liver IRI by Suzuki’s histological grading. Mean±SD, **P<0.001; ##P<0.001.

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Figure 8 Expand