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Table 1.

Primer sequences and expected amplification products.

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Figure 1.

Changes in relative weight, TNF-R1 expression and morphology in endocrine glands from T. cruzi-infected mice.

The relative weight of the adrenal glands in C57BL/6 acutely infected with T. cruzi, at day 17 pi (panel A) was calculated as follows: [adrenal weight (mg)/body weight (g)]. Panel B shows western blots for the protein expression of TNF-R1 at day 17 pi. Results are expressed as mean ± SEM from 3–5 mice/group. A representative experiment from 2 independent series is shown. # p<0.05 vs. uninfected counterparts. Panel C. The cortex consists of three morphologically distinct zones. The region immediately beneath the capsule is called zone glomerulosa and constitutes about 1/5 or less of the entire thickness, this zone was only increased in Tc-WT mice. The mid portion of the cortex, called zone fasciculata, comprises about 50–65% of the thickness of the cortex in uninfected mice (WT and TNF-R1−/− mice), with Tc-WT and Tc-TNF-R1−/− mice showing a diffuse hyperplasia (the broad arrow indicates the thickness with an arbitrary vertical line). The most inner portion of the cortex, called zone reticularis, comprises about 20–25% of the cortex from uninfected mice, showing a hyperplasia in Tc-TNF-R1−/− mice. H&E stain, magnification 40 X. Scale bar: 20 µm. WT (wild type) and TNF-R1−/− (TNF receptor deficient) control mice; Tc-WT and Tc-TNF-R1−/− infected counterparts.

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Table 2.

Cytokine levels in plasma of mice undergoing acute T. cruzi infection.

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Table 3.

Morphometric analysis of adrenal sections of mice undergoing acute T. cruzi infection.

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Figure 2.

Hormonal variations in the HPA axis from T. cruzi-infected mice.

Plasma levels of corticosterone (µg/dl, panel A) and ACTH (pg/ml, panel B) in C57BL/6 mice acutely infected with T. cruzi. Results are expressed as mean ± SEM from 3–5 mice/group. A representative experiment from 2 independent series is shown. WT (wild type) and TNF-R1−/− (TNF receptor deficient) control mice; Tc-WT and Tc-TNF-R1−/− infected counterparts. *p<0.05 vs. Tc-WT; #p<0.05 vs. uninfected counterparts.

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Figure 3.

Activation of NF-κB and AP-1 in adrenal gland during T. cruzi infection.

Levels of NF-κB activity (OD 450/650, panel A) and activation of AP-1 (panel B) in nuclear extracts of adrenal glands from C57BL/6 mice acutely infected with T. cruzi. NF-κB activity assay was assessed by colorimetric determination of p50 subunit in all experimental groups. AP-1 activation was tested using direct measurement of c-jun by Western blot. Results are shown as mean ± SEM from 3–5 mice/group/day. A representative experiment from two similar rounds is shown. WT (wild type) and TNF-R1−/− (TNF receptor deficient) control mice; Tc-WT and Tc-TNF-R1−/− infected counterparts. #p<0.05 vs. uninfected counterparts; &p<0.05 vs. WT mice.

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Figure 4.

Activation of MAPK pathways in adrenal glands from mice acutely infected with T.cruzi .

Blots correspond to P-p38, P-ERK and P-JNK. Blots were stripped and revealed with an anti-β-actin (panel A). Bars represent the densitometry considering WT and TNF-R1−/− as 100% (panel B). Expression is relative to β-actin. Results are expressed as mean ± SEM from 3–5 mice/group. Data correspond to one round of two representative and independent experimental rounds. WT (wild type) and TNF-R1−/− (TNF receptor deficient) control mice; Tc-WT and Tc-TNF-R1−/− infected counterparts. *p<0.05 vs. Tc-WT; #p<0.05 vs. uninfected counterparts.

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Figure 5.

Steroidogenic enzymes gene expression in adrenal glands from T. cruzi infection in absence of TNF-R1.

Expression profiles of genes involved in GC synthesis/activation enzymes, StAR (panel A), CYP11A1 (panel B), CYP11B1 (panel C), 11β-HSD1 (panel D) and 11β-HSD2 (panel E) in adrenal glands from C57BL/6 or TNF-R1−/− mice acutely infected with T. cruzi. Total mRNA was purified and analyzed by Real-Time-PCR using nonsaturating conditions. The mRNA expression levels were then normalized to RPL13. Results are shown as mean ± SEM from 3–4 mice/group/day. A representative experiment from 3 independent series is shown. WT (wild type) and TNF-R1−/− (TNF receptor deficient) control mice; Tc-WT and Tc-TNF-R1−/− infected counterparts. *p<0.05 vs. Tc-WT; #p<0.05 vs. uninfected counterparts and &p<0.05 vs. WT.

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Figure 6.

Gene expression of pro-inflammatory cytokines in adrenal glands from T. cruzi-infected mice.

Total RNA was extracted and the mRNA levels of TNF-α (panel A), IL-6 (panel B) and IL-1β (panel C) were assessed by quantitative PCR (normalized to RPL13). Results are shown as mean ± SEM from 3–4 mice/group/day. A representative experiment from 3 independent series is shown. WT (wild type) and TNF-R1−/− (TNF receptor deficient) control mice; Tc-WT and Tc- TNF-R1−/− infected counterparts. *p<0.05 vs. Tc-WT; #p<0.05 vs. uninfected counterparts.

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Figure 7.

Expression of TNF receptor type 2 in adrenal gland from T. cruzi- infected mice.

Panel A shows frozen sections of the adrenal gland labeled with anti-TNF-R2 antibody. Confocal microscopy shows a decreased in TNF-R2 contents in cortex of the adrenal gland in Tc-WT and Tc-TNF-R1−/− (more pronounced in Tc-TNF-R1−/− group) after 17 days of infection. The small box is a representative control staining in which an unrelated primary antibody was applied. Panel B shows graphs corresponding to the relative quantification analysis of cortical TNF-R2 expression of 5–6 microscopic fields of adrenal gland from WT (wild type) and TNF-R1−/− (TNF receptor deficient) control mice and infected counterparts (Tc-WT and Tc-TNF-R1−/−). Panel C shows the gene expression of TNF-R2 in adrenal glands from T. cruzi-infected mice. Results are expressed as mean ± SEM. A representative experiment from 2 independent series is shown. The scale bar represents 50 µm (C: cortex; M: medulla). Original magnifications 200X. * p<0.05 vs. Tc-WT; # p<0.05 vs. uninfected counterparts.

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Figure 8.

Detection of T. cruzi in the adrenal gland.

Panel A shows frozen sections of the adrenal gland labeled with anti–T. cruzi antiserum. Immunostaining analysis demonstrated alike deposition of parasite antigens in the adrenal glands from Tc-WT and Tc-TNF-R1−/− after 17 days of infection. The small boxes represent a control staining in which an unrelated primary antibody was applied. Panel B corresponds to the relative quantification analysis of cortical T.cruzi-labeling of 5–6 microscopic fields of adrenal glands from WT (wild type) and TNF-R1−/− (TNF receptor-1 deficient) control mice and their infected counterparts (Tc-WT and Tc-TNF-R1−/−). Panel C shows the quantitative expression of T. cruzi kDNA in adrenal glands from T. cruzi-infected mice. Results are expressed as mean ± SEM. A representative experiment from 2 independent series is shown. The scale bar represents 50 µm. Original magnifications 200X.

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