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Figure 1.

In situ ELISA assay workflow.

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Table 1.

Marine seaweeds selected for antiviral screening, location of collection and cytotoxicity activity in Huh7.5 cells.

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Figure 2.

In situ ELISA optimization.

(A) Effect of different blocking buffers (skim milk and FBS) and fixatives (methanol:acetone and paraformaldehyde followed by triton X-100) for the assay. (B) Cells were seeded in a range of concentration and the influence for the assay performance was evaluated. (C) The ideal MOI for all DENV serotypes was evaluated 48 hours post-infection and (D) 72 hours post-infection. Values represent mean ± SD of three independent experiments performed in triplicate.

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Figure 3.

In situ ELISA assay validation.

(A) Correlation between the in situ ELISA and the foci-forming assay, where Huh7.5 was infected with DENV-1, DENV-2, DENV-3 and DENV-4 at a range of MOI (4–0.01). After 72 hours of infection the supernatant was used for the foci-forming assay and the cells submitted to the ELISA assay. Data from one representative experiment, analyzed using Pearson correlation test. (B) Comparison of the DRC and the IC50 for IFN-α 2A and DENV-4 infection, obtained with the in situ ELISA, foci-forming assay and a commercial NS1 antigen capture ELISA assay. Mean ± SD of three independent experiments, analyzed by sigmoidal dose-response curve (variable slope), the dashed line represents the detection limit of the foci-forming assay.

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Figure 4.

Seaweed extracts antiviral screening.

(A) Huh7.5 were seeded in a 96-well plate and infected with DENV-1, (B) DENV-2 and (C) DENV-3 with an MOI of 4, and (D) DENV-4 with an MOI of 0.1. Interferon-α 2A (100 IU/ml) was used as a positive control and after 72 hours post-infection the ELISA was performed. Data were analyzed using one-way ANOVA followed by Tukey test. Values are mean ± SD of three independent experiments. *p<0.05.

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Figure 5.

DENV infection inhibition for different strains/serotypes.

Huh7.5 was seeded in a 96-well plate and infected with each strain with a MOI of 4 for 1 h30 min, then the seaweed extracts were added and after 72 hours the ELISA was performed. Data were analyzed using one-way ANOVA followed by Dunnett test. Values are mean ± SD of three independent experiments,*p<0.05.

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Figure 6.

Time of addition studies with seaweed extracts A1, A3, A8 and A12.

(A) Huh 7.5 cells were infected with DENV-4 with an MOI of 0.1 following each treatment (before the infection, −1.5 h; during the infection, 0 h; and after the infection, +1.5 h) at the MNTD, the cells were submitted to the ELISA assay (B) and the supernatant for the foci-forming assay. The dashed line represents the detection limit of the foci-forming assay.

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Figure 7.

Dose response curve for the seaweed extracts A1, A3, A8 and A12.

(A) Huh 7.5 cells were infected with DENV-4 and treated during the infection in a range of concentration for each seaweed extract. (B) CC50, IC50 and SI were calculated. Mean ± SD of three independent experiments, analyzed by sigmoidal dose response curve (variable slope).

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Figure 8.

Effect of the seaweed extracts A1, A3, A8 and A12 on DENV-4 entry process.

(A) Binding inhibition assay. (B) Internalization inhibition assay. Data were analyzed using one-way ANOVA followed by Dunnett test. Values are mean ± SD of three independent experiments,*p<0.05.

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