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Figure 1.

Expression of CD4 on cells co-cultured with or without Notch signaling.

(A) CB and (B) Adult HSCs were maintained in OP9Deltapos (□) or OP9Deltaneg (▪) co-cultures for 28 days with weekly analysis of CD4+ content by flow cytometry. Both CB and adult - derived HSCs expressed CD4 without Notch signaling. Data presented as mean ± SEM (n = 10 for OP9Deltapos and n = 4 for OP9Deltaneg). (C–F) Representative flow cytometry plots showing CD4 and CD14 expression on cells grown in co-culture for 28 days. (C) CB in OP9Deltaneg, (D) CB in OP9Deltapos, (E) adult in OP9Deltaneg, (F) adult in OP9Deltapos.

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Figure 2.

Growth of adult and CB cells on OP9Deltapos co-culture.

(A) Representative dot plot (Adult cells, 84 days, 7 days DAPT treatment) showing expression of CD3 and CD4 within the DP population. (B) Plot of CD3 generation by CB (▪) and adult (▵) cells in OP9Deltapos co-culture. (C) Cell number in OP9Deltapos CB (▪) and adult (▵) co-culture (mean ± SEM, n = 9).

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Figure 3.

Notch signal interruption promotes CD4 and DP cell differentiation.

CB and adult HSCs were grown in OP9Deltapos co-culture, then Notch signalling was removed for 7 days by transfer to OP9Deltaneg co-culture or addition of DAPT. Cells were analysed by flow cytometry for CD4 and CD8 expression. (A) Representative dot plots for adult cells co-cultured for 70 days. (B–E) Adult (B, D) or CB (C, E) HSCs were grown in OP9Deltapos co-culture (▪) and then either transferred to OP9Deltaneg co-culture (○) or incubated with DAPT (▵) for 7 days. Data points show the percentage of CD4+ ISP (B, C) or CD4+ CD8+ DP (D, E) cells, n≥3, mean+SEM, * p<0.05 for DAPT vs OP9Deltapos (t-test), ** p<0.05 for OP9Deltaneg vs OP9Deltapos (t-test). (F) CD14 (▪) expression plotted against DP expression (▴) for adult cells in OP9Deltapos co-cultures after Notch signalling was removed for 7 days. (G) Cell numbers during DAPT treatment. Paired comparisons of six control and DAPT – treated cultures, showing all HSC-derived cells (left) and DP cells (right).

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Figure 4.

Gene expression differences in HSCs co-cultured on OP9Delta cells with or without Notch signalling.

Real-time RT-PCR results from cord blood (CB) or adult HSCs co-cultured with either control OP9Deltaneg (C) cells, or OP9Deltapos (+N) cells. Results are depicted as mean expression ratio +SEM relative to housekeeping gene HPRT (n = 3–4 for CB; n = 2–4 for adults). * p<0.05 compared with ‘d28 +N’ for CB and adult respectively, unpaired 2-tail t-test. d = day. PTCRA = pre-T cell antigen receptor alpha, GATA3 = GATA binding protein 3, IKZF1 = IKAROS family zinc finger 1, TCF7 = Transcription Factor 7 (T-cell specific, HMG-box), HES1 = Hairy and enhancer of split 1, DTX1 = Deltex homolog 1, MAML1 = Mastermind-like 1, CDKN1B = Cyclin-dependent kinase inhibitor 1B.

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Figure 5.

Effect of Notch signal removal (DAPT) on mRNA expression of CB and adult HSCs undergoing T lineage differentiation.

Real-time RT-PCR results from cells grown in OP9Deltapos co-culture with Notch inhibitor (DAPT) added for 7 days before harvesting. Timepoints: CB (cord blood) early = 42 days, CB late = 56 days, A (adult) early = 42 days, A late = 70 days. Data are presented as mean fold change in mRNA expression + SEM relative to untreated control cultures (OP9Deltapos), with baseline set at 1.0 (n = 3 for CB; n = 2–4 for adults). * p<0.05 compared with the control, paired 2-tail t-test.

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Table 1.

PCR primers.

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Table 1 Expand