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Figure 1.

Recruitment of skin-derived DC in mice developing Amela-melanomas is impaired, but their expression of CCR7 is unaltered.

(A) Ratio of MigDC versus ResDC in Amela-TDLN or corresponding LN from control mice. (B) % TRITC-labeled cells among MigDC in Amela-TDLN or corresponding LN from control mice 20 hrs after TRITC-skin painting (see Methods). (C-F) Analysis of DC subpopulations in LNs from control (C, E) or Amela-bearing (D, F) mice after TRITC-painting. (C-D) After gating on CD11c+NK1.1CD25CD45R cells, TRITC-staining is shown versus SSC. TRITC and TRITC+ cells were analyzed for expression of CD11c versus MHCII. ResDC and MigDC were found within TRITC cells; only MigDC were found within TRITC+ cells. Only MigDC expressed CD207 corresponding to Langerhans cells and CD207+ Dermal DC. (E–F) Histograms of CCR7 expression for the different DC subpopulations indicated in C and D, respectively, show low, intermediate and high fluorescence values (mean), respectively, for ResDC, CD207MigDC and CD207+MigDC.

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Figure 1 Expand

Figure 2.

TL localization in TDLN of mice developing Amela-melanomas is impaired.

(A) CFSE-labeled CD8 TL from B10.D2 mice (white) were adoptively transferred into control, Mela- and Amela-bearing recipient mice. 15 hrs later their LN (A) were stained with anti-B220-PE-Texas red (red) and anti-CD3-APC (blue) and spleens (B) were stained with anti-B220-APC (blue) (upper pictures) or anti-MOMA-1-biotyl/streptavidin-Alexa647 (blue) (lower pictures) and analyzed by confocal microscopy (see Methods). The graph in (A) represents the number of CFSE+ cells per section of LN. The graph in (B) represents the number of CFSE+ cells per splenic white pulp. Number of white pulps analyzed per mouse was > or = 2. Data are from 8 (control), 3 (Mela-bearing) and 4 (Amela-bearing) mice. Statistics are as described in Methods.

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Figure 2 Expand

Figure 3.

Lymphoid organization in developing Amela-melanomas is disrupted.

Analysis of LN (A) and spleen (B) from control, Mela- and Amela-bearing mice. (A) In LN, B- and T-zones were identified with anti-B220-PE-Texas red (red) and anti-CD3-FITC (green), respectively. Graphs represent the density of B (left) and T (right) cells, determined by pixel quantification. For tumor-bearing mice, TDLN are shown. Data are from 7 (control), 4 (Mela-bearing) and 4 (Amela-bearing) mice. (B) Splenic B-zones, T-zones and marginal zones were identified with anti-B220-APC (blue), anti-CD3-FITC (green), and anti-MOMA-1/goat anti-rat-Alexa647 (red), respectively. The white pulp areas are delineated by MOMA-1 expression. One graph (left) shows the size of white pulp areas in the different mice (each symbol represents a white pulp area). Density of B (middle graph) and T (right graph) cells were determined by pixel quantification. Data are from 6 (control), 4 (Mela-bearing) and 9 (Amela-bearing) mice.

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Figure 3 Expand

Figure 4.

Decrease in splenic lymphoid chemokines in mice developing Amela-melanomas.

Spleen sections from control, Mela- and Amela-bearing mice stained in (A) with anti-B220-APC (blue) to detect the B-zone, ER-TR7/chicken anti-rat-Alexa488 (green) to detect the FRC network and anti-CCL21/donkey anti-goat-Alexa546 (red). (B) Expression of ccl21a encoding Ccl21-Ser, ccl19 and cxcl13 relative to tbp transcripts measured by QRT-PCR in total spleens of control, Mela- or Amela- bearing mice. Means (+/− s.e.m.) for independent samples (N = 10; 8; 14 for ccl21a; N = 5; 4; 8 for ccl19; N = 3; 3; 9 for cxcl13) from spleens of control, Mela- or Amela- bearing mice, respectively, A is representative of 5 mice per group.

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Figure 5.

Loss of gp38+ FRC-associated CCL21 in spleens and TDLN of mice developing Amela-melanomas.

In (A) spleen sections were stained with ER-TR7/chicken anti-rat-Alexa647 (blue), anti-gp38/donkey anti-goat-Alexa488 (green) and anti-CCL21-biotyl/SA-Alexa546 (red). In (B) LN sections were stained with anti-B220-Alexa647 (blue), anti-gp38/donkey anti-goat-Alexa546 (red) and anti-CCL21-biotyl/SA-Alexa488 (green). A and B are representative of 3 mice per group. Lower images show magnifications of the fields highlighted in the upper images and show stainings with each of the antibodies separately and merged, as indicated.

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Figure 6.

Splenic architecture is disrupted, with loss of contractile fibroblasts, in mice developing Amela-melanomas.

Spleens from control, Mela-, and Amela-bearing mice stained in (A) with ER-TR7/chicken anti-ratAlexa488 (left), anti-laminin/donkey anti-rabbit-Alexa488 (middle), and anti-collagen IV/donkey anti-rabbit-Alexa488 (rigth) to detect stromal networks, and in (B) with anti-B220-FITC (green), anti-Desmin/donkey anti-rabbit-Alexa555 (red). Representative of 5 mice per group.

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Figure 7.

Gr1+CD11b+ cells recruited in SLO interact with the stroma in mice developing Amela-melanomas.

LN (A) or spleen (B) sections from control, Mela- and Amela-bearing mice were stained with anti-Gr1-PE (red) and anti-Lyve-1-biotyl/streptavidin-FITC (green) to detect LN medullas or for MOMA-1 (green) to delineate splenic white pulp areas, respectively. LN (C) or spleen (D) sections from Amela-bearing mice were stained with anti-Gr1-APC (green) and anti-ER-TR7/chicken anti-rat-Alexa647 (blue). Magnification showing a cluster of Gr1+ cells on the stromal network is shown.

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Figure 7 Expand