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Figure 1.

Immunohistochemical validation of genes identified in microarray experiments.

A representative ETS abnormal skeletal muscle fiber is shown A. Prohibitin 2, B. Mitochondrial DNA Polymerase Gamma, C. P53 Up-regulated Mediator of Apoptosis, D. Cytochrome C Oxidase activity, E. Succinate Dehydrogenase activity.

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Figure 2.

Immunohistochemical validation of signal transduction/transcriptional activation identified by gene expression profiling.

Activation of AMP kinase and peroxisome proliferator activated receptor pathways in response to deletion mutation accumulation. A. CD36/Fatty acid Translocase, a pparα regulated gene, B. No Primary antibody control, C. Peroxisome proliferator-activated receptor gamma co-activator 1, D. Activated AMP Kinase, E. Inhibited Acetyl-CoA Carboxylase F. Peroxisome proliferator-activated receptor alpha.

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Figure 3.

Effect of β-GPA administration on mitochondrial DNA abundance in vivo.

A. Mitochondrial genome content of the Vastus medialis muscle following β-GPA treatment was measured using real-time PCR. B. Electron transport system abnormalities are more abundant in rats treated with the AMP kinase activator β-guanidinopropionic acid. Each Vastus lateralis skeletal muscle was stained for cytochrome C oxidase and succinate dehydrogenase every 60 microns for one millimeter. Regions of skeletal muscle fibers lacking COX activity and hyper-reactive for SDH (the ETS abnormal phenotype) were counted.

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Figure 4.

Model of positive feedback loop in ETS abnormal fibers.

Signal transduction pathways detect mitochondrial dysfunction and drive transcriptional activation leading to up-regulation of mitochondrial DNA replication and subsequent deletion mutation accumulation. Genes in green were up-regulated in ETS abnormal fibers. Proteins in blue were found to be up-regulated by immunohistochemistry in ETS abnormal fibers. Proteins in purple were detected by both assays. wt: wild-type mitochondrial genomes, Δ deletion mutation containing mitochondrial genomes.

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