Spatial and seasonal variation in macrozoobenthic density, biomass and community composition in a major tropical intertidal area, the Bijagós Archipelago, West-Africa

• Ana Pinto Coelho ,

  Contributed equally to this work with: Ana Pinto Coelho, Mohamed Henriques

  Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Validation, Visualization, Writing – original draft

  * E-mail: apcoelho@ua.pt

  Affiliation Centre for Environmental and Marine Studies (CESAM), Department of Biology, University of Aveiro, Aveiro, Portugal

  https://orcid.org/0000-0002-3337-0310

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• Mohamed Henriques ,

  Contributed equally to this work with: Ana Pinto Coelho, Mohamed Henriques

  Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Validation, Visualization, Writing – original draft

  Affiliations Centre for Environmental and Marine Studies (CESAM), Department of Animal Biology, Faculdade de Ciências da Universidade de Lisboa, Lisbon, Portugal, Conservation Ecology Group, Groningen Institute for Evolutionary Life Sciences, University of Groningen, Groningen, The Netherlands

  https://orcid.org/0000-0002-3061-5423

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• Afonso Duarte Rocha,

  Roles Investigation, Methodology, Writing – review & editing

  Affiliations Centre for Environmental and Marine Studies (CESAM), Department of Biology, University of Aveiro, Aveiro, Portugal, Conservation Biology Research Group, Department of Anatomy, Cell Biology and Zoology, University of Extremadura, Badajoz, Spain

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• João Paulino,

  Roles Data curation, Investigation, Methodology, Writing – review & editing

  Affiliation Centre for Environmental and Marine Studies (CESAM), Department of Animal Biology, Faculdade de Ciências da Universidade de Lisboa, Lisbon, Portugal

  https://orcid.org/0000-0003-0779-1960

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• Loran Kleine Schaars,

  Roles Data curation, Investigation, Methodology, Resources, Writing – review & editing

  Affiliation Department of Coastal Systems, NIOZ Royal Netherlands Institute for Sea Research, Texel, The Netherlands

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• Catarina Ramos,

  Roles Data curation, Investigation, Writing – review & editing

  Affiliation Centre for Environmental and Marine Studies (CESAM), Department of Biology, University of Aveiro, Aveiro, Portugal

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• Aissa Regalla de Barros,

  Roles Funding acquisition, Project administration, Resources, Writing – review & editing

  Affiliation Instituto da Biodiversidade e das Áreas Protegidas Dr. Alfredo Simão da Silva (IBAP), Bissau, Guiné-Bissau

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• Teresa Catry,

  Roles Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – review & editing

  Affiliation Centre for Environmental and Marine Studies (CESAM), Department of Animal Biology, Faculdade de Ciências da Universidade de Lisboa, Lisbon, Portugal

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• José Pedro Granadeiro,

  Roles Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – review & editing

  Affiliation Centre for Environmental and Marine Studies (CESAM), Department of Animal Biology, Faculdade de Ciências da Universidade de Lisboa, Lisbon, Portugal

  https://orcid.org/0000-0002-7207-3474

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• Theunis Piersma,

  Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – review & editing

  Affiliations Conservation Ecology Group, Groningen Institute for Evolutionary Life Sciences, University of Groningen, Groningen, The Netherlands, Department of Coastal Systems, NIOZ Royal Netherlands Institute for Sea Research, Texel, The Netherlands

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• José Augusto Alves

  Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – review & editing

  Affiliations Centre for Environmental and Marine Studies (CESAM), Department of Biology, University of Aveiro, Aveiro, Portugal, South Iceland Research Centre, University of Iceland, Laugarvatn, Iceland

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