Reproductive parameters and cub survival of brown bears in the Rusha area of the Shiretoko Peninsula, Hokkaido, Japan

• Michito Shimozuru ,

  * E-mail: shimozuru@vetmed.hokudai.ac.jp

  Affiliation Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

  http://orcid.org/0000-0002-3281-8208

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• Masami Yamanaka,

  Affiliation Shiretoko Museum, Shari, Japan

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• Masanao Nakanishi,

  Affiliation Shiretoko Nature Foundation, Shari, Japan

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• Jun Moriwaki,

  Affiliation Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

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• Fumihiko Mori,

  Affiliation Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

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• Masakatsu Tsujino,

  Affiliation Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

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• Yuri Shirane,

  Affiliation Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

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• Tsuyoshi Ishinazaka,

  Affiliation Shiretoko Nature Foundation, Shari, Japan

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• Shinsuke Kasai,

  Affiliation Shiretoko Nature Foundation, Shari, Japan

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• Takane Nose,

  Affiliation Shiretoko Nature Foundation, Shari, Japan

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• Yasushi Masuda,

  Affiliation Shiretoko Nature Foundation, Shari, Japan

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• Toshio Tsubota

  Affiliation Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

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