Are Eyes a Mirror of the Soul? What Eye Wrinkles Reveal about a Horse’s Emotional State

• Sara Hintze ,

  * E-mail: sara.hintze@vetsuisse.unibe.ch

  Affiliations Division of Animal Welfare, Vetsuisse Faculty, University of Bern, Bern, Switzerland, Agroscope, Swiss National Stud Farm, Avenches, Switzerland

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• Samantha Smith,

  Affiliation Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

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• Antonia Patt,

  Affiliation Department of Animal and Avian Sciences, University of Maryland, College Park, Maryland, United States of America

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• Iris Bachmann,

  Affiliation Agroscope, Swiss National Stud Farm, Avenches, Switzerland

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• Hanno Würbel

  Affiliation Division of Animal Welfare, Vetsuisse Faculty, University of Bern, Bern, Switzerland

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