Diet and Macronutrient Optimization in Wild Ursids: A Comparison of Grizzly Bears with Sympatric and Allopatric Black Bears

• Cecily M. Costello ,

  * E-mail: cecostello@mt.gov

  Current address: Montana Fish, Wildlife, and Parks, Kalispell, Montana, United States of America

  Affiliation College of Forestry and Conservation, University of Montana, Missoula, Montana, United States of America

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• Steven L. Cain,

  Current address: Grand Teton National Park Foundation, Moose, Wyoming, United States of America

  Affiliation Grand Teton National Park, Moose, Wyoming, United States of America

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• Shannon Pils,

  Current address: Shoshone National Forest, Cody, Wyoming, United States of America

  Affiliation U. S. Geological Survey, Northern Rocky Mountain Science Center, Interagency Grizzly Bear Study Team, Bozeman, Montana, United States of America

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• Leslie Frattaroli,

  Current address: National Park Service, National Capital Region, Washington, DC, United States of America

  Affiliation Grand Teton National Park, Moose, Wyoming, United States of America

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• Mark A. Haroldson,

  Affiliation U. S. Geological Survey, Northern Rocky Mountain Science Center, Interagency Grizzly Bear Study Team, Bozeman, Montana, United States of America

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• Frank T. van Manen

  Affiliation U. S. Geological Survey, Northern Rocky Mountain Science Center, Interagency Grizzly Bear Study Team, Bozeman, Montana, United States of America

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