Barley HvHMA1 Is a Heavy Metal Pump Involved in Mobilizing Organellar Zn and Cu and Plays a Role in Metal Loading into Grains

• Maria Dalgaard Mikkelsen,

  Affiliations Centre for Membrane Pumps in Cells and Disease (PUMPKIN), Danish National Research Foundation, Frederiksberg, Denmark, Department of Plant and Environmental Sciences, University of Copenhagen, Frederiksberg, Denmark

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• Pai Pedas,

  Affiliation Department of Plant and Environmental Sciences, University of Copenhagen, Frederiksberg, Denmark

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• Michaela Schiller,

  Affiliation Department of Plant and Environmental Sciences, University of Copenhagen, Frederiksberg, Denmark

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• Eva Vincze,

  Affiliation Department of Molecular Biology and Genetics, Research Centre Flakkebjerg, Aarhus University, Slagelse, Denmark

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• Rebecca F. Mills,

  Affiliation Centre for Biological Sciences, University of Southampton, Southampton, Hampshire, United Kingdom

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• Søren Borg,

  Affiliation Department of Molecular Biology and Genetics, Research Centre Flakkebjerg, Aarhus University, Slagelse, Denmark

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• Annette Møller,

  Affiliations Centre for Membrane Pumps in Cells and Disease (PUMPKIN), Danish National Research Foundation, Frederiksberg, Denmark, Department of Plant and Environmental Sciences, University of Copenhagen, Frederiksberg, Denmark

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• Jan K. Schjoerring,

  Affiliation Department of Plant and Environmental Sciences, University of Copenhagen, Frederiksberg, Denmark

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• Lorraine E. Williams,

  Affiliation Centre for Biological Sciences, University of Southampton, Southampton, Hampshire, United Kingdom

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• Lone Baekgaard,

  Affiliations Centre for Membrane Pumps in Cells and Disease (PUMPKIN), Danish National Research Foundation, Frederiksberg, Denmark, Department of Plant and Environmental Sciences, University of Copenhagen, Frederiksberg, Denmark

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• Preben Bach Holm,

  Affiliation Department of Molecular Biology and Genetics, Research Centre Flakkebjerg, Aarhus University, Slagelse, Denmark

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• Michael G. Palmgren

  * E-mail: palmgren@life.ku.dk

  Affiliations Centre for Membrane Pumps in Cells and Disease (PUMPKIN), Danish National Research Foundation, Frederiksberg, Denmark, Department of Plant and Environmental Sciences, University of Copenhagen, Frederiksberg, Denmark

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