Pathogen Webs in Collapsing Honey Bee Colonies

• R. Scott Cornman,

  Affiliation Bee Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

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• David R. Tarpy,

  Affiliation Department of Entomology, North Carolina State University, Raleigh, North Carolina, United States of America

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• Yanping Chen,

  Affiliation Bee Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

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• Lacey Jeffreys,

  Affiliation Department of Entomology, North Carolina State University, Raleigh, North Carolina, United States of America

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• Dawn Lopez,

  Affiliation Bee Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

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• Jeffery S. Pettis,

  Affiliation Bee Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

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• Dennis vanEngelsdorp,

  Affiliation Department of Entomology, University of Maryland, College Park, Maryland, United States of America

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• Jay D. Evans

  * E-mail: Jay.evans@ars.usda.gov

  Affiliation Bee Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland, United States of America

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