An Ancient Origin for the Enigmatic Flat-Headed Frogs (Bombinatoridae: Barbourula) from the Islands of Southeast Asia

• David C. Blackburn ,

  * E-mail: david.c.blackburn@gmail.com

  Affiliation Biodiversity Institute, and Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas, United States of America

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• David P. Bickford,

  Affiliation National University of Singapore, Singapore, Singapore

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• Arvin C. Diesmos,

  Affiliation National Museum of the Philippines, Ermita, Manila, Philippines

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• Djoko T. Iskandar,

  Affiliation Institut Teknologi Bandung, Bandung, Java, Indonesia

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• Rafe M. Brown

  Affiliation Biodiversity Institute, and Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas, United States of America

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