About the Authors

Maddalena M. Bolognesi

Contributed equally to this work with: Maddalena M. Bolognesi, Lorenzo Dall’Olio, Gastone Castellani

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Software, Visualization, Writing – original draft

Affiliations Istituto di Bioimmagini e Sistemi Biologici Complessi (IBSBC) – CNR Via F.lli Cervi, Segrate, Italy, NBFC, National Biodiversity Future Center, Palermo, Italy

Lorenzo Dall’Olio

Contributed equally to this work with: Maddalena M. Bolognesi, Lorenzo Dall’Olio, Gastone Castellani

Roles Conceptualization, Formal analysis, Investigation, Methodology, Software, Visualization, Writing – original draft

Affiliation Laboratorio di Data Science and Bioinformatics, IRCCS Istituto delle Scienze Neurologiche di Bologna – AUSL BO Ospedale Bellaria, Bologna, Italy

Giulio Eugenio Mandelli

Roles Formal analysis, Resources, Validation, Writing – original draft

Affiliations Pathology Unit, Department of Molecular and Translational Medicine-DMMT, University of Brescia, Brescia, Italy, Pathology Unit, ASST Spedali Civili Di Brescia, Brescia, Italy

Luisa Lorenzi

Roles Data curation, Investigation, Resources

Affiliations Pathology Unit, Department of Molecular and Translational Medicine-DMMT, University of Brescia, Brescia, Italy, Pathology Unit, ASST Spedali Civili Di Brescia, Brescia, Italy

Francesca M. Bosisio

Roles Funding acquisition, Resources, Writing – original draft

Affiliations The Leuven Institute for Single-cell Omics (LISCO), KULeuven, Leuven, Belgium, Translational Cell and Tissue Research Unit, Department of imaging and pathology, KULeuven, Leuven, Belgium, Pathology Department, University Hospital of Leuven, Leuven, Belgium

Ann M. Haberman

Roles Investigation, Resources, Writing – original draft

Affiliations Department of Immunobiology, Yale University, New Haven, Connecticut, United States of America, Department of Laboratory Medicine, Yale University, New Haven, Connecticut, United States of America

Govind Bhagat

Roles Resources, Writing – original draft

Current address: Pathology, Saint Louis University School of Medicine, Saint Louis, MO, USA

Affiliation Pathology, Columbia University Irving Medical Center and New York Presbyterian Hospital, NewYork, New York, United States of America

Simone Borghesi

Roles Formal analysis, Methodology, Software, Writing – original draft

Affiliation Department of Mathematics and Applications, University of Milano Bicocca, Milan, Italy

Mario Faretta

Roles Data curation, Investigation, Methodology, Software, Validation

Affiliation Department of Experimental Oncology, European Institute of Oncology IRCCS, Milan, Italy

Gastone Castellani

Contributed equally to this work with: Maddalena M. Bolognesi, Lorenzo Dall’Olio, Gastone Castellani

Roles Funding acquisition, Methodology, Supervision

Affiliation Department of Experimental, Diagnostic and Specialty Medicine, University of Bologna, Bologna, Italy

Giorgio Cattoretti

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

giorgio.cattoretti@unimib.it

Affiliation Pathology, Department of Medicine and Surgery, Universitá di Milano-Bicocca, Monza, Michigan, Italy

Competing Interests

The authors have declared that no competing interests exist.