About the Authors

Yume Okamoto

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Resources, Validation, Visualization, Writing – original draft

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Madoka Hattori

Roles Funding acquisition, Investigation, Validation, Writing – review & editing

Affiliations Wildlife Research Center, Kyoto University, Kyoto, Japan, Kyoto City University of Arts, Kyoto, Japan

Miho Inoue-Murayama

Roles Conceptualization, Funding acquisition, Project administration, Resources, Supervision, Writing – review & editing

murayama.miho.5n@kyoto-u.ac.jp

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.