About the Authors

Jung Yun Kim

Roles Conceptualization, Formal analysis, Writing – original draft

Affiliations Department of Biotechnology, College of Life Sciences and Biotechnology, Korea University, Seoul, Republic of Korea, Institute of Animal Molecular Biotechnology, Korea University, Seoul, Republic of Korea

Nayoung Hong

Roles Investigation, Writing – review & editing

Affiliations Department of Biotechnology, College of Life Sciences and Biotechnology, Korea University, Seoul, Republic of Korea, Institute of Animal Molecular Biotechnology, Korea University, Seoul, Republic of Korea

Seok Won Ham

Roles Investigation, Writing – review & editing

Affiliation MEDIFIC Inc., Hwaseong-si, Gyeonggi-do, Republic of Korea

Sehyeon Park

Roles Investigation, Writing – review & editing

Affiliations Department of Biotechnology, College of Life Sciences and Biotechnology, Korea University, Seoul, Republic of Korea, Institute of Animal Molecular Biotechnology, Korea University, Seoul, Republic of Korea

Sunyoung Seo

Roles Investigation, Writing – review & editing

Affiliations Department of Biotechnology, College of Life Sciences and Biotechnology, Korea University, Seoul, Republic of Korea, Institute of Animal Molecular Biotechnology, Korea University, Seoul, Republic of Korea

Hyunggee Kim

Roles Conceptualization, Supervision, Writing – review & editing

hg-kim@korea.ac.kr

Affiliations Department of Biotechnology, College of Life Sciences and Biotechnology, Korea University, Seoul, Republic of Korea, Institute of Animal Molecular Biotechnology, Korea University, Seoul, Republic of Korea

Competing Interests

H.K. is the founder, majority shareholder, and external director of MEDIFIC Inc. S.W.H. was affiliated with MEDIFIC, Inc. The authors declare no conflict of interest. This does not alter our adherence to PLOS ONE policies on sharing data and materials.