About the Authors

Qing Yang

Roles Conceptualization, Data curation, Formal analysis, Methodology, Writing – original draft

s1362029@ed.ac.uk

Affiliation Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Cathy M. Dwyer

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliations Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom, Department of Animal Behaviour and Welfare, Scotland’s Rural College (SRUC), Edinburgh, United Kingdom

Belinda Vigors

Roles Conceptualization, Supervision, Writing – review & editing

Affiliation Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Ruqian Zhao

Roles Supervision

Affiliation College of Veterinary Science, Nanjing Agricultural University, Nanjing, China

Fritha M. Langford

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliations Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom, School of Natural and Environmental Science, Newcastle University, Newcastle upon Tyne, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.